De Herbario Berolinense Notulae No. 21
H.J.M. SIPMAN

Lichenotheca Latinoamericana a museo botanico 
berolinensi edita, fasciculum primum

Abstract

Sipman, H.J.M.: Lichenotheca Latinoamericana a museo 
botanico berolinensi edita, fasciculum primum [Lichenotheca 
Latinoamericana edited by the Botanical Museum Berlin-
Dahlem, first fascicle]. - Willdenowia 19: 543-551. 1990. - 
ISSN 0511-9618.

Label information is presented on 50 lichen specimens from 
Brazil, Colombia, Costa Rica, French Guyana, Guyana, 
Mexico and Surinam, which form the first fascicle of this 
series of exsiccata. Substantial range extensions documented 
by these specimens include the first finding in the Neotropis 
of Chroodiscus mirificus and the first record from Colombia 
of Cystocoleus ebeneus. Heterodermia flavosquamosa from 
Guyana is described as a species new to science.

This new series of exsiccata intends to contribute to the 
investigation of the lichenized fungi occuring in Latin 
America. It will be issued in 25 sets, which are distributed to 
the following herbaria: B, C, CBG, COL, COLO, GZU, LD, 
M, NY, PRE, TNS, U, UPS, US, VEN. The remaining sets 
will preferentially be distributed to latinamerican herbaria.
The first fascicle contains material of an unpublished species 
of Heterodermia, as well as records of several major range 
extensions. Therefore a list with label information, together 
with a description of the new species, is published in a 
recognized botanical journal rather than as a special leaflet.
With Lichenotheca Latinoamericana a tradition of the lichen 
herbarium of the Botanical Museum Berlin-Dahlem is 
resumed, which was started by G. Follmann in 1968 when 
issuing "Lichenes Exsiccati Selecti a Museo Botanico 
Berolinensi Editi". From this series four fascicles were 
issued: fasc. I, II and III in 1968 and fasc. IV in 1970 (see 
Willdenowia 4: 383-390, 4: 391-397, 5: 15-21 and 6: 17-23). 
From Fasc. V onward the series was issued under the title 
"Lichenes Exsiccati Selecti a Museo Historiae Naturalis 
Casselensi Editi" (see Philippia 2: 13-21) from the 
Naturkundemuseum Kassel, where Follmann had accepted a 
position.


1. Biatorella conspersa (Fée) Vainio
det. H. Sipman, 1989. First record for the Guianas.
GUYANA, Upper Mazaruni District, along trail from 
Kamarang river to Pwipwi Mountain, c. 10 km N of 
Waramadan, 60 45' W, 5 57' N, in rocky savannah, c. 800 m.
leg. H. Sipman & A. Aptroot 19470. d.d. 28 February 1985

2. Buellia efflorescens Müller Arg.
det. A. Aptroot, 1985. Material cited in Fl. Guianas E1: 12 
(1988 ("1987")).
GUYANA, East Demerara District, along western dike at 
mouth of Mahaica river, cultivated coastal area, 57 55' W, 
6 41-43' N, on trunk of Cocos nucifera, 0 m.
leg. H. Sipman & A. Aptroot 19509. d.d. 5 March 1985

3. Chroodiscus mirificus (Kremp.) R. Sant.
det. H. Sipman, 1988. First record for the Neotropis.
GUYANA, Upper Mazaruni District, Kamarang, in the 
village, 60 37' W, 5 52' N, on leaves of Mangifera tree in 
garden, c. 500 m.
leg. H. Sipman & A. Aptroot 18149. d.d. 26 February 1985

4. Cladina confusa (Sant.) Follm. & Ahti
det. H. Sipman, 1977. TLC: usnic and perlatolic acids.
COLOMBIA, Depart. Boyacá, along road Ráquira - La 
Candelaria, km 4.5, terrestrial on rocky NE-facing slope with 
Andropogon spp., 2450 m.
leg. A. M. Cleef, H. García-B. & R. Jaramillo-M. 3473. d.d. 6 
May 1972

5. Cladina densissima Ahti
det. Teuvo Ahti, 1987. TLC: usnic and fumarprotocetraric 
acids.
GUYANA, Upper Mazaruni District, along trail from 
Kamarang river to Pui-Pui Mountain, c. 10 km N of 
Waramadan, 60 45' W, 5 57' N, in rocky savannah, c. 800 m.
leg. H. Sipman & A. Aptroot 19334. d.d. 28 February 1985

6. Cladina sprucei (Ahti) Ahti 
det. Teuvo Ahti, 1987. TLC: atranorin and fumarprotocetraric 
acid.
GUYANA, Upper Mazaruni District, mount Latipu, c. 8 km 
N of Kamarang, 60 38' W, 5 57' N, on white sand on open 
place in scrub on the summit plateau, c. 800 m.
leg. H. Sipman & A. Aptroot 19129. d.d. 25 February 1985

7. Cladonia ceratophylla (Sw.) Sprengel
det. H. Sipman, l986. TLC: fumarprotocetraric acid.
SURINAM, Wilhelmina mountains, in tall forest on slope 
near camp Km 14 on line to Lucie River, at c. 2 m heigth on 
treetrunk, c. 400 m.
leg. J.P. Schulz 10453. d.d. 25 July l963

8. Cladonia corallifera (Kunze) Nyl.
det. Teuvo Ahti, l987. TLC: usnic, didymic and thamnolic 
acids.
GUYANA, East Demerara District, Timehri, Dakara Creek, 
Thomson's farm, 58 15' W, 6 29' N, terrestrial in savannah-
forest remnant on white sand, c. 10 m.
leg. H. Sipman & A. Aptroot 19500. d.d. 2 February l985

9. Cladonia didyma (Fée) Vainio var. vulcanica (Zoll. & 
Mor.) Vainio
det. Teuvo Ahti, 1987. TLC: didymic, barbatic and thamnolic 
acids.
GUYANA, Upper Mazaruni District, along trail from 
Kamarang river to Pui-Pui Mountain, c. 10 km N of 
Waramadan, 60 45' W, 5 57' N, in rocky savannah, c. 800 m.
leg. H. Sipman & A. Aptroot 19497. d.d. 28 February 1985

10. Cladonia furcata (Hudson) Schrader 
det. H. Sipman, 1979. TLC: fumarprotocetraric acid.
COLOMBIA, Depart. Meta, Cerro Nevado del Sumapaz, 
Páramo de Sumapaz, on humid rocks in paramo on NW-
slope, near limestone cave where the river disappears, 3550 
m. 
leg. A.M. Cleef 7619. d.d. 10 January 1973

11. Cladonia signata (Eschw.) Vainio 
det. Teuvo Ahti, 1987. TLC: fumarprotocetraric acid.
GUYANA, Upper Mazaruni District, along trail from 
Kamarang river to Pui-Pui Mountain, c. 10 km N of 
Waramadan, 60 45' W, 5 57' N, in rocky savannah, c. 800 m.
leg. H. Sipman & A. Aptroot 19327. d.d. 28 February 1985

12. Cladonia signata (Eschw.) Vainio
det. Teuvo Ahti, 1987. TLC: atranorin and fumarprotocetraric 
acid.
GUYANA, Upper Mazaruni District. along trail from 
Kamarang river to Pui-Pui Mountain, c. 10 km N of 
Waramadan, 60 45' W, 5 57' N, in rocky savannah, c. 800 m.
leg. H. Sipman & A. Aptroot 19331. d.d. 28 February 1985

13. Cladonia squamosa (Scop.) Hoffm. var. squamosa
det. H. Sipman, 1986. TLC: squamatic acid.
COSTA RICA, prov. San José, Cordillera de Talamanca, 
Cerro de la Muerte, along Carretera Interamericana, on 
decaying log in open place in humid subparamo scrub, 3325 
m.
leg. A.M. Cleef & L.A. Fournier 10215. d.d. 21. August l973

14. Coccocarpia palmicola (Sprengel) Arvidsson & D. 
Galloway c. apoth.
det. A. Aptroot, 1985.
FRENCH GUYANA, Cayenne, Montagne du Mahury, c. 10 
km SE of Cayenne, forest relict on vulcanic soil, 52 17' W, 
4 53' N, on tree exposed to seawind, 100 m.
leg. A. Aptroot 15464. d.d. March 1985

15. Coenogonium linkii Ehrenb.
det. A. Aptroot, 1985. Apothecia on lower surface.
FRENCH GUYANA, Saül, primary forest near Roche Bateau, 
53 10' W, 3 38' N, on sapling, 200-300 m.
leg. A. Aptroot 15386. d.d. March l985

16. Collema callibotrys Tuck. var. coccophyllizum (Zahlbr.) 
Degel.
det. H. Sipman, l986.
COSTA RICA, prov. Cartago, area Recreativa Prusia, c. 3 km 
N of Tierra Blanca, near Mirador, 84 53' W, 9 57' N, 
epiphyte in disturbed, shrubby, montane forest, 2700 m.
leg. H. Sipman 20495. d.d. 15 March l985

17. Cystocoleus ebeneus (Dillw.) Thwaites
det. H. Sipman, 1985. First record for Colombia.
COLOMBIA, Depart. Cundinamarca, Municipio Supatá, Alto 
El Tablazo, along trail from radar station to Supatá, in 
subparamo scrub, on perpendicular sandstone rockface, 3400 
m.
leg. J. Aguirre C. & H. Sipman 5228. d.d. 11 September l984

18. Dictyonema glabratum (Sprengel) D. Hawksw.
det. H. Sipman, l987.
GUYANA, Upper Mazaruni District, along trail from 
Kamarang River to Pui-Pui Mountain, c. 5 km N of 
Waramadan, 60 46' W, 5 54' N, in savannah bush, c. 650 m.
leg. H. Sipman & A. Aptroot 19245. d.d. 27 February - 1 
March l985

19. Erioderma chilense Mont.
det. H. Sipman, l986.
COSTA RICA, prov. San José, Cerro de la Muerte, along 
Interamerican Highway, c. 2 m tall scrub with rock outcrops, 
E of Cerro Buenavista, 83 46' W, 9 33' N, epiphyte, 3400 m.
leg. H. Sipman 20958. d.d. 23 March l985

20. Erioderma sorediatum D. Galloway & Jørgensen
det. H. Sipman, l986.
COSTA RICA, Prov. San José, Cerro de la Muerte, along 
Interamerican Highway, c. 2 m tall scrub with rock outcrops, 
E of Cerro Buenavista, 83 46' W, 9 33' N, epiphyte, 3400 m.
leg. H. Sipman 20959. d.d. 23 March l985

21. Erioderma verruculosum Vainio
det. H. Sipman, l986.
COSTA RICA, prov. San José, Cerro de la Muerte, along 
Interamerican Highway, c. 2 m tall scrub with rock outcrops, 
E of Cerro Buenavista, 83 46' W, 9 33' N, epiphyte, 3400 m.
leg. H. Sipman 20957. d.d. 23 March l985

22. Everniastrum vexans (Zahlbr.) Hale ex Sipman
det. H. Sipman, l986. TLC: atranorin, protolichesterinic and 
salazinic acids.
COSTA RICA, prov. Cartago, SW-slope of Volcán Turrialba, 
N of Santa Cruz, Fila Bonilla, along Camino El Trazado, 
83 43' W, 10 01' N, pastureland with relic trees of cloud 
forest, 2100 m.
leg. H. Sipman 20423. d.d. 14 March l985

23. Heterodermia erinacea (Ach.) W.A.Weber
det. H. Sipman, 1989. TLC: atranorin and zeorin.
MEXICO, Baja California, along route 1, near km 131 from 
Ensenada to San Quintin, 2 km S of Colonet, 116 13' W, 
31 04' N, on shrub branchlets in c. 1 m tall desert scrub on 
hill with volcanic rock outcrops, coastal scrub community 
with e.g. Agave shawii, Ambroisia chenopodifolia, 
Machaerocereus gummosus, 100 m.
leg. H. Sipman 24768. d.d. 3 January l989

24. Heterodermia flavosquamosa Aptroot & Sipman - 
ISOTYPUS 
det. A. Aptroot & H. Sipman.
GUYANA, Upper Mazaruni District, along trail from 
Kamarang River to Pui-Pui Mountain, c. 5 km N of 
Waramadang, 60 46' W, 5 54' N, on exposed rock in 
savannah bush, c. 650 m.
leg. H. Sipman & A. Aptroot 19233. d.d. 27-28 February l985




Heterodermia flavosquamosa Aptroot & Sipman, sp. nov.
Typus: Sipman & Aptroot 19233 (Holo- B; iso- distributed in 
Lichenotheca Latinoamericana 1: 24).

Thallus appressus vel apicibus ascendentibus, laciniis 
linearibus, pagina inferiore decorticata arachnoidea 
flavescente sed centrum versus denudata obscurascente, 
margine thalli et apotheciorum squamulis ecorticatis albis 
fragilibus applanatis vel teretibus proviso.

Thallus appressed, yellowish grey, up to 10 cm diam. Lobes 
flat, 0.4-0.6 mm wide, with 0.7-1.4 mm long internodes, at 
tips flat or ascending. Soredia and isidia absent. Squamules 
numerous, mostly ecorticate, flattened or more or less terete, 
mainly marginal on flabellate lobe tips or on small lateral 
lobes, sometimes also on low superficial warts, fragile, white 
or slightly bluegrey, often with yellow pigmentation on 
underside, 0.2-0.4 mm high, c. 0.1 mm thick, often 
resembling sorediose isidia because they are easily 
fragmented. Upper surface corticate, shiny. Medulla white. 
Lower surface ecorticate, near the lobe tips with arachnoid 
yellow-pigmented cover, in older parts denudated and 
darkening or violet grey. Rhizinae absent on lower side but 
present as cilia on the margins of the lobes and sometimes on 
the squamules, simple or with a few thin perpendicular 
branchlets, black, 2-4 mm long, c. 0.1 mm thick. 
Chemistry: atranorin, an unidentified anthraquinone, zeorin 
and several unknown terpenoids present. Reactions: cortex K+ 
y, C-, P-, UV-; medulla K+ y, C-, P-, UV-; yellow pigment K+ 
red, C+ red, P+ red, also in tap water + red, UV-. 
Apothecia sessile, becoming substipitate, 2-5 mm diam. Disc 
darkbrown, concave to flat. Margin squamulose, the 
squamules with yellow side directed towards disc, incurved 
over the disc. Hymenium 100-130 µm thick. Spores of 
Polyblastidium-type, 25-30 x 10-15 µm, 8 per ascus.

The new species is known from the Guianas and Venezuela. It 
seems to be restricted to the hinterland and has been found on 
mossy rock in savannas and mountain areas.
The type has been cited in Aptroot, Fl. Guianas E1: 27 (1988 
("1987")) as H. corallophora (Taylor) Skorepa. Examination 
of the holotype (FH-Taylor) and an isotype (BM) proved, 
however, that H. Corallophora differs by the presence of 
genuine isidia and the absence of yellow pigment on the lower 
side.
Heterodermia flavosquamosa looks much like a squamulose 
counterpart of H. flabellata (Fée) Awasthi (primary species) 
and H. obscurata (Nyl.) Trevisan (sorediate). However, the 
pigment shows different colour reactions.

Additional material examined: SURINAM: Wilhelmin 
Gebergte, near Julian Top, Irwin et al. 55182a (NY); 
Voltzberg, summit area, J. & P.A. Florschütz 4566 (U). 
VENEZUELA: Estado Bolivar, distrito Cedeño, vicinity of 
Panare village, Boom 6459 & Grillo (NY).

25. Heterodermia leucomelos (L.) Poelt subsp. leucomelos
det. A. Aptroot, 1985. Material cited in Fl. Guianas E1: 30 
(1988 ("1987")); only record for the Guianas. TLC: atranorin, 
terpenoids and salazinic acid.
GUYANA, Upper Mazaruni District, along trail from 
Kamarang River to Pui-Pui Mountain, c. 10 km N of 
Waramadang, 60 45' W, 5 57' N, on branchlets in rocky 
savannah, c. 800 m.
leg. H. Sipman & A. Aptroot 19284. d.d. 28 February l985

26. Hyperphyscia adglutinata (Flörke) Mayrh. & Poelt
det. A. Aptroot, 1985. Material cited in Fl. Guianas E1: 34 
(1988 ("1987")).
GUYANA, West Berbice district, Onverwach, cultivated 
coastal area, c. 57 35' W, 6 25' N, epiphyte on Avicennia 
treetrunk in grazed remnants of old mangrove, 0 m.
leg. H. Sipman & A. Aptroot 19609. d.d. 9 March l985

27. Leptogium laceroides Bouly de Lesdain
det. H. Sipman, l986.
COSTA RICA, prov. San José, Cerro de la Muerte, along 
Interamerican Highway, c. 2 m tall scrub with rock outcrops, 
E of Cerro Buenavista, 83 46' W, 9 33' N, epiphyte, 3400 m.
leg. H. Sipman 20956. d.d. 23 March l985

28. Leptogium phyllocarpum (Pers.) Mont.
det. H. Sipman, l980.
COSTA RICA, prov. Limón, road along the Caribbean coast 
from Limón to Cahuita, between Rio Estrella and R. 
Margarita, on shrubs on sand along the beach, in closed, c. 6 
m tall bush, 0 m.
leg. H. Sipman 12306. d.d. 9 January l979

29. Microtheliopsis uleana Müller Arg.
det. H. Sipman, l988. First record for the Guianas.
GUYANA, Upper Mazaruni District, along trail from 
Kamarang River to Pui-Pui Mountain, at landing site, c. 5 km 
NW of Waramadang, 60 47' W, 5 48' N, foliicolous in 
undergrowth of c. 25 m tall forest, c. 600 m.
leg. H. Sipman & A. Aptroot 19225c. d.d. 27-28 February 
l985

30. Parmotrema dilatatum (Vainio) Hale
det. R. van Aubel, 1985. First record for the Guianas. TLC: 
atranorin, usnic, echinocarpic and protocetraric acids.
GUYANA, Upper Mazaruni District, Kamarang, in the 
village, 60 37' W, 5 52' N, on free-standing tree on river 
bank, c. 500 m.
leg. H. Sipman & A. Aptroot 18194. d.d. 26 February l985

31. Peltigera vainioi Gyelnik
det. O. Vitikainen, 1987. First record for Colombia.
COLOMBIA, Depart. Risaralda, Municipio Santa Rosa de 
Cabal, W-slope of Cordillera Central, along track from 
Termales de Santa Rosa to Termales Antiguos, on humid, 
loamy bank, 2000 m.
leg. J. Aguirre C. & H. Sipman 5570. d.d. 19 September 1984

32. Pertusaria acroscyphoides Sipman in Willdenowia 16: 
281 (1987) - ISOTYPUS 
det. H. Sipman. TLC: protocetraric acid.
COLOMBIA, Depart. Boyacá, Paramos NW of Belén, 
between Laguna Grande and El Púlpito, common lichen in 
vegetation with Plantago monticola, Draba, Racomitrium, 
Thamnolia and Bidens sp., 3830 m. 
leg. A.M. Cleef 9781. d.d. 6 May 1973

33. Phylloporis phyllogena (Müller Arg.) Clements
det. A. Vezda, l985.
BRAZIL, Estado São Paulo, between Mogi das Cruzes and 
Bertioga, Serra do Garrãozinho, S of Manuel Ferreira along 
road SP-39, on leaf in undergrowth in c. 15 m tall forest on 
slope, c. 850 m.
leg. H. Sipman 12706. d.d. 30 July l979

34. Physma byrsaeum (Ach.) Tuck.
det. A. Aptroot, 1985.
FRENCH GUYANA, Montagne du Mahury, forest relict on 
vulcanic soil c. 10 km SE of Cayenne, 52 17' W, 4 53' N, on 
tree exposed to seawind, 100 m.
leg. A. Aptroot 15461. d.d. March 1985

35. Porina epiphylla (Fée) Fée
det. A. Vezda, l985.
BRAZIL, Estado São Paulo, between Mogi das Cruzes and 
Bertioga, Serra do Garrãozinho, S of Manuel Ferreira along 
road SP-39, on leaf in undergrowth in c. 15 m tall forest on 
slope, c. 850 m.
leg. H. Sipman 12707c. d.d. 30 July l979

36. Pyxine cocoes (Sw.) Nyl.
det. A. Aptroot, 1985. Material cited in Fl. Guianas E1: 45 
(1988 ("1987")). TLC: lichexanthone.
GUYANA, West Berbice District, Onverwacht, cultivated 
coastal plain, c. 57 35' W, 6 25' N, epiphyte on Avicennia 
treetrunk in grazed remnants of old mangrove, 0 m.
leg. H. Sipman & A. Aptroot 19604. d.d. 9 March l985

37. Ramalina ceruchis Ach.
det. H. Sipman, 1989. TLC: traces of div. terpenoids, 
bourgeanic acid?
MEXICO, Baja California, Cabo San Quintin, along Pacific 
Ocean front NW of Bahía Falsa, 116 02' W, 30 28' N, on 
shrub branches in c. 1 m tall desert scrub at the seashore, on 
W-side of low volcano with basalt rocks, coastal scrub 
community with e.g. Euphorbia, Jatropha, c. 10 m.
leg. H. Sipman 24865. d.d. 4 January l989

38. Ramalina crinita Tuck.
det. H. Sipman, 1989. TLC: traces of terpenoids.
MEXICO, Baja California, near km 45 on road from San 
Quintin to Parador Punta Prieta, c. 10 km N of El Rosaria, in 
Cañon del Rosario, 115 46' W, 30 08' N, on branch of shrub 
in c. 1 m tall desert scrub on hill of post-batholithic marine 
sedimentary (Cretaceous) rocks, coastal scrub community with 
e.g. Aesculus, Jatropha, Lycium, 90 m.
leg. H. Sipman 24903. d.d. 5 January l989

39. Ramalina josecuervoi Rundell & Bowler
det. H. Sipman, 1989. TLC: salazinic acid.
MEXICO, Baja California, at seashore 25 km N of Guerrero 
Negro, between Laguna Manuela and Punta Santo Domingo, 
114 06' W, 28 14' N, on rock in gravelly plain with post-
batholithic basalt outcrops and sand with scattered desert 
scrub of Euphorbia misera above rocky seashore (Southern 
Viscaíno Region of the Sonoran Desert (Fouquieria diguetii, 
Jatropha, Yucca, etc.)), c. 10 m.
leg. H. Sipman 24989. d.d. 7 January l989

40. Ramalina leptocarpha Tuck.
det. H. Sipman, 1989. TLC: traces of terpenoids, bourgeanic 
acid?
MEXICO, Baja California, along route 1, near km 131 from 
Ensenada to San Quintin, 2 km S of Colonet, 116 13' W, 
31 04' N, on shrub branchlets in c. 1 m tall desert scrub on 
hill with volcanic rock outcrops, coastal scrub community 
with e.g. Agave shawii, Ambroisia chenopodifolia, 
Machaerocereus gummosus, 100 m.
leg. H. Sipman 24757. d.d. 3 January l989

41. Ramalina subleptocarpha Rundell & Bowler
det. H. Sipman, 1989. TLC: usnic acid, traces of terpenoids, 
bourgeanic acid?
MEXICO, Baja California, near km 170 on road from Santa 
Rosalia to Guerrero Negro, c. 3 km on sideroad to Punta 
Eugenia, W of Vizcaíno junction, 113 25' W, 27 39' N, on 
branches of shrubs in desert scrub on sandy plain (Southern 
Viscaíno Region of the Sonoran Desert (Fouquieria diguetii, 
Jatropha, Pachycereus pringlei, etc.)), 60 m.
leg. H. Sipman 25094. d.d. 9 January l989

42. Rinodina sipmanii Aptroot in Proc. Kon. Ned. Akad. 
Wetensch. C 90: 241 (1987) - ISOTYPE 
det. A. Aptroot. Material cited in Fl. Guianas E1: 53 (1988 
("1987")).
GUYANA, East Demerara District, along western dike at 
mouth of Mahaica river, cultivated coastal plain, 57 55' W, 
6 41-43' N, on tree in village, 0 m.
leg. H. Sipman & A. Aptroot 19505. d.d. 5 March l985

43. Sphaerophorus melanocarpus (Sw.) DC.
det. H. Sipman, 1975. TLC: perlatolic and stictic acids.
COLOMBIA, Depart. Arauca, Sierra Nevada del Cocuy, plain 
of San José in valley of El Playón, terrestrial lichen on 
Chusquea cushions in humid paramo dominated by 
Chusquea, with Oritrophium limnophilum subsp. mutisianum, 
Alchemilla paludicola and Breutelia, 3680 m. 
leg. A. M. Cleef 10095. d.d. 9 July 1973

44. Stereocaulon atlanticum (Lamb) Lamb
det. H. Sipman, 1985. TLC: atranorin, perlatolic (trace) and 
protocetraric acids.
COLOMBIA, Depart. Risaralda, Municipio Santa Rosa de 
Cabal, W-slope of Cordillera Central, c. 500 m S of Finca La 
Sierra, on vertical rockface of escarpment in clearings in 
subparamo woodland, 3750 m.
leg. J. Aguirre C. & H. Sipman 5419. d.d. 16 September l984

45. Stereocaulon strictum Th. Fr. var. strictum
det. H. Sipman, 1985. TLC: atranorin, porphyrilic acid.
COLOMBIA, Depart. Cauca, Municipio, Puracé, E-side of 
Cordillera Central, along road La Plata - Puracé, 2 km E of 
Termales de San Juan, on bank of road in humid high-andean 
woodland, 3000 m.
leg. J. Aguirre C. & H. Sipman 5871. d.d. 4 October l984

46. Stereocaulon vesuvianum Pers. var. nodulosum (Wallr.) 
Lamb
det. T. Boekhout, 1978. TLC: atranorin, norstictic, stictic, 
cryptostictic and menegazziaic acids.
COLOMBIA, Depart. Caldas, Nevado del Ruíz, superparamo 
zone, 3 km NW of Refugio, terrestrial lichen forming 
cushions on black sand on NW-facing slope, 4300 m.
leg. A. M. Cleef & H. 't Hart 2555. d.d. 19 March l972

47. Sticta humboldtii J.D. Hooker
det. H. Sipman, 1985.
COLOMBIA, Depart. Risaralda, Municipio Santa Rosa de 
Cabal, W-slope of Cordillera Central, c. 500 m S of Finca La 
Sierra, epiphyte in remnant of subparamo woodland, 3750 m.
leg. J. Aguirre C. & H. Sipman 5458. d.d. 16 September l984

48. Sticta weigelii (Isert) Ach.
det. A. Aptroot, l987.
FRENCH GUYANA, Saül, primary forest near Boeuf Mort, 
53 12' W, 3 38' N, on buttresses of Ceiba pentandra, 200-300 
m.
leg. A. Aptroot 15342. d.d. March l985

49. Strigula nemathora Mont.
det. H. Sipman, l988.
GUYANA, Upper Mazaruni District, along trail from 
Kamarang River to Pui-Pui Mountain, at landing site, c. 5 km 
NW of Waramadan, 60 47' W, 5 48' N, foliicolous in 
undergrowth in c. 25 m tall forest, c. 600 m.
leg. H. Sipman & A. Aptroot 19229b. d.d. 27-28 February 
l985

50. Thamnolia vermicularis (Sw.) Schaerer 
det. H. Sipman, 1975. TLC: thamnolic acid.
COLOMBIA, Depart. Boyacá, Páramo de la Sarna between 
Sogamoso and Vado Hondo, 5 km NE of Laguna de Tota, 
terrestrial lichen associated with Lorenzochloa erectifolia in 
very wind-exposed paramo, 3550 m.
leg. A.M. Cleef 9203. d.d. 29 March 1973


Address of the author:
Dr. H.J.M. Sipman, Botanischer Garten und Botanisches 
Museum Berlin-Dahlem, Königin-Luise-Str. 6-8, D-14195 
Berlin.