De Herbario Berolinensi Notulae No. 27 
ROSEMARIE GEBAUER 
 
The type specimens of African Loranthaceae described by others than 
Engler and kept in the general herbarium at Berlin-Dahlem (B) 
 
Abstract 
Gebauer, R.: The type specimens of African Loranthaceae described by 
others than Engler and kept in the general herbarium at Berlin-Dahlem (B). 
- Willdenowia 23: 291-304. 1993. - ISSN 0511-9618. 
About 100 type specimens of African Loranthaceae used for description of 
new taxa by authors other than Engler are represented in the general 
herbarium at Berlin-Dahlem. The types are listed alphabetically with their 
locality data, collecting dates, status (as far as known) and their accepted 
names. An index to the collectors is included. 
 
[Note: The equals sign for nomenclatural synonyms is coded as "=#"]. 
 
Introduction 
Besides c. 300 type specimens of Engler's Loranthus taxa (Gebauer 1993) 
the general herbarium at Berlin-Dahlem holds about 100 type specimens of 
African Loranthaceae described by other authors, especially by Sprague. 
The original material used by Engler is in Berlin and duplicates are in Kew 
whereas the greater part of the original material used by Sprague is in Kew 
and duplicates are in Berlin. 
Among the listed types are altogether about 20 individual collections which 
were used by different authors for the descriptions of different new taxa. 
The arrangement of information is the same as in Gebauer (1993). The data 
are cited from the labels, but complementary data from the protologues are 
added enclosed in square brackets when thought helpful. 
 
List of the African Loranthaceae described by others than Engler and their 
type specimens in B 
 
1. Korthasella madagascarica Danser in Bull. Jard. Bot. Buitenzorg, ser. 3, 
14: 126 (1937). 
– Madacascar, bassin de réception de la Mananara, affluent du Mandrare 
pentes occidentales des montagnes entre l'Andohahela et l'Elakelaka, au 
Vatazo (Sud d'Imonty), 900-950 m, 1.-2. 1934, Humbert 14060 (type, fide 
Balle 1964: 111). 
 
2. Loranthus albizziae De Wild. var. rogersii Sprague in Thiselton Dyer, Fl. 
Trop. Afr. 6(1): 289 (1910). 
– Zambia, near Ndola, Rogers 8318 (isotype, fide Polhill in sched.). 
= Phragmanthera cornetii (Dewèvre) Polhill & Wiens (fide Polhill in 
sched.). 
 
3. Loranthus alveatus Sprague in Thiselton Dyer, Fl. Trop. Afr. 6(1): 315 
(1910). 
– Tanzania, Sansibarküste, Doda, 27. 6. 1893, Holst 2946 (holotype, fide 
Polhill in litt.). - Also syntype of Loranthus dregei Eckl. & Zeyh. f. 
subcuneifolius Engl. 
#= Erianthemum alveatum (Sprague) Danser (fide Danser 1933: 53). 
 
4. Loranthus brachyanthus Peter in Repert. Spec. Nov. Regni Veg. Beih. 
40(2): 177, Anh.: 15 (1932). 
– Tanzania, Usaramo, Dar-es-Salaam, Bagomoyolim, im Pori, 1. 9. 1926, 
Peter 44752 (type, fide Polhill in sched.). 
= Englerina triplinervia (Baker & Sprague) Polhill & Wiens (fide Polhill in 
sched.). 
 
5. Loranthus brazzavillensis De Wild. in Bull. Soc. Roy. Bot. Belgique 
39(2): 102 (1900). 
– Congo, Brazzaville, 2. 1891, Demeuse 119 (isotype). - Also syntype of 
Loranthus crassicaulis Engl. 
= Tapinanthus constrictiflorus (Engl.) Danser (fide Polhill in litt. 10/1991). 
 
6. Loranthus buchneri Engler var. gossweileri Sprague in Thiselton Dyer, 
Fl. Trop. Afr. 6(1): 358 (1910). 
– Angola, Loanda, 1903, Gossweiler 149 (isosyntype, fide Polhill in litt.). - 
Also holotype of Loranthus gracilis Engl. & K. Krause 
= Tapinanthus buchneri (Engl.) Danser (fide Danser 1933: 109). 
– Angola, Loanda, 1903, Gossweiler 473 (isosyntype, fide Polhill in litt.). - 
Also holotype of Loranthus obtusilobus Engl. & K. Krause. 
= Tapinanthus buchneri (Engl.) Danser (fide Danser 1933: 109). 
 
7. Loranthus buntingii Sprague in Bull. Misc. Inform. Kew 1916: 178 
(1916). 
– Liberia, [Mount Barclay], Bunting 165 (isotype). 
#= Tapinanthus buntingii (Sprague) Danser (fide Danser 1933: 109). 
 
8. Loranthus bussei Sprague in Thiselton Dyer, Fl. Trop. Afr. 6(1): 299 
(1910). 
– Tanzania, Namgaru-Ufer beim Dt. Namgaru, 18. 6. 1903, Busse 2949 (2 
isolectotypes, chosen by Polhill & Wiens ined.). 
#= Oliverella bussei (Sprague) Polhill & Wiens (fide Polhill & Wiens 1992: 
169). 
 
9. Loranthus carsonii Baker & Sprague in Thiselton Dyer, Fl. Trop. Afr. 
6(1): 376 (1910). 
– Zambia, Tanganyika-Plateau, [Urungu], Fwambo, Carson s.n. (isotype, 
fide Polhill in sched.). 
= Agelanthus pungu (De Wild.) Polhill & Wiens (fide Polhill in sched.). 
 
10. Loranthus ceciliae N. E. Br. in Bull. Misc. Inform. Kew 1906: 168 
(1906). 
– Zimbabwe, near Buluwayo [Waterworks], Cecil 96 (2 isotypes, fide 
Polhill in sched.). 
= Agelanthus pungu (De Wild.) Polhill & Wiens (fide Polhill in sched.). 
 
11. Loranthus cistoides Welw. ex Engl. var. longiflorus Schinz in Bull. 
Herb. Boissier 4, Append. 3: 52 (1896). 
–  Namibia, Grootfontein, [Südost-Ondonga], 10. 1886, Schinz 294 
(isosyntype, fide Polhill in sched.). 
= Phragmanthera glaucocarpa (Peyr.) Balle (fide Balle 1968: 167). 
 
12. Loranthus deltae Baker & Sprague in Thiselton Dyer, Fl. Trop. Afr. 
6(1): 319 (1910). 
– Mozambique, Luabo, mouths of Zambesi [and Kongone Rivers], Kirk s.n. 
(fragment of type, fide Polhill in sched.). 
#= Agelanthus deltae (Baker & Sprague) Polhill & Wiens (fide Polhill & 
Wiens 1992: 163). 
 
13. Loranthus dregei Eckl. & Zeyh., Enum. Pl. Afric. Austral. 358 (1837). 
– South Africa, [Bothasberg prope flumen 'Vischrivier' (Albany)], [Eckl. & 
Zeyh. 2284] (type, fide Polhill in sched.). - Also syntype of Loranthus 
dregei Eckl. & C. Zeyh. f. subcuneifolius Engl. 
#= Erianthemum dregei (Eckl. & Zeyh.) Tiegh. (fide Polhill in sched.). - 
There are two other specimens in B which are probably additional type 
specimens; however, their labels lack collection dates. 
 
14. Loranthus dregei Eckl. & Zeyh. var. foliaceus Sprague in Thiselton 
Dyer, Fl. Trop. Afr. 6(1): 314 (1910). 
– Ethiopia, Harar, 28. 4. 1900, Ellenbeck 797 (type). 
= Erianthemum dregei (Eckl. & C. Zeyh.) Tiegh. (fide Polhill in sched.). 
– Ethiopia, Walenzo, 9. 7. 1900, Ellenbeck 1303 (type). 
= Erianthemum dregei (Eckl. & Zeyh.) Tiegh. (fide Polhill in sched.). 
 
15. Loranthus dregei Eckl. & Zeyh. var. kerenicus Sprague in Thiselton 
Dyer, Fl. Trop. Afr. 6(1): 312 (1910). 
– Eritrea, Bogos, 6.1870, Beccari 82 (isolectotype, fide Polhill in sched.). - 
Also syntype of Loranthus dregei Eckl. & Zeyh. f. obtusifolius Engl. 
= Erianthemum dregei (Eckl. & Zeyh.) Tiegh. (fide Polhill in sched.). 
– Eritrea, Keren, 9. 1872, Hildebrandt 517 (2 types). - Also syntypes of 
Loranthus dregei Eckl. & Zeyh. f. obtusifolius Engl. 
= Erianthemum dregei (Eckl. & Zeyh.) Tiegh. (fide Polhill in sched.). 
– Ethiopia/Sudan, Gallabat, Matamma, 7. 1865, Schweinfurth 2193 (type). - 
Also syntype of Loranthus dregei Eckl. & Zeyh. f. obtusifolius Engl. 
= Erianthemum dregei (Eckl. & Zeyh.) Tiegh. (fide Polhill in sched.). 
 
16. Loranthus dregei Eckl. & Zeyh. var. kilimanjaricus Sprague in Thiselton 
Dyer, Fl. Trop. Afr. 6(1): 314 (1910). 
– Tanzania, Kilimandscharo, Marangu, Marktplatz, 1500 m, 22. 2. 1894, 
Volkens 1934 (2 isotypes, fide Polhill in sched.). 
= Erianthemum dregei (Eckl. & Zeyh.) Tiegh. (fide Polhill in sched.). 
 
17. Loranthus dregei Eckl. & Zeyh. var. longipes Sprague in Thiselton 
Dyer, Fl. Trop. Afr. 6(1): 314 (1910). 
– Tanzania, bei Kwa Kihingi (Ungoni), 2. 1. 1901, Busse 764 (holotype, 
fide Polhill in sched.). 
= Erianthemum lindense (Sprague) Danser (fide Polhill in sched).  

18. Loranthus dregei Eckl. & Zeyh. var. nyasicus Sprague in Thiselton 
Dyer, Fl. Trop. Afr. 6(1): 313 (1910). 
– Tanzania, Uluguru-Gebirge, 1000 m, 21. 11. 1898, Goetze 162 (syntype). 
= Erianthemum schelei (Engl.) Tiegh. (fide Polhill in sched.). 
– Tanzania, Ussangu, nördl. Kinga-Berge [Kinga (Livingstone) 
Mountains.], Mingi-Berg, 2100 m [in Engler's handwriting on the label of 
the isosyntype: "Rondo Plateau"], 6. 6. 1899, Goetze 1014 (syntype, 
isosyntype). 
= Erianthemum dregei (Eckl. & Zeyh.) Tiegh. (fide Polhill in sched.). 
 
19. Loranthus dregei Eckl. & Zeyh. var. ovatus Sprague in Thiselton Dyer, 
Fl. Trop. Afr. 6(1): 314 (1910). 
– Tanzania, Amani, 8. 1908, Braun 1970 (2 types). 
= Erianthemum dregei (Eckl. & Zeyh.) Tiegh. (fide Polhill in sched.). 
– Tanzania, Ost-Usambara, Amani, 17. 9. 1902, Engler 707 (2 types). 
= Erianthemum dregei (Eckl. & Zeyh.) Tiegh. (fide Polhill in sched.). 
 
20. Loranthus elegantissimus Schinz in Bull. Herb. Boissier 4, Append. 3: 
52 (1896). 
– Namibia, Amboland, Oombale, 3. 1896, Schinz 291 (isosyntype, fide 
Polhill in sched.). 
= Oncocalyx welwitschii (Engl.) Polhill & Wiens (fide Polhill in sched.). 
 
21. Loranthus engleri Hiern, Cat. Afr. Pl. Welw. 1:  930 (1900). 
– Angola, Pungo Andongo, 11. 1856, Welwitsch 4849 (isotype, fide Polhill 
in litt.). - Also isotype of Loranthus emarginatus Engl. 
#= Phragmanthera engleri (Hiern) Polhill & Wiens (fide Polhill & Wiens 
1992: 171). 
 
22. Loranthus eucalyptoides Peter in Repert. Spec. Nov. Regni Veg. Beih. 
40(2): 178, Anh.: 16 (1932), non DC., nom illegit. 
– Tanzania, Landschaft Meru, Urwald beim Forsthause am Meru-Berg, 
1800 m ü. M., 24./26. 2. 1914, Peter 1888 (holotype, fide Polhill in sched.). 
= Englerina woodfordioides (Schweinf.) Balle (fide Polhill & Wiens 1992: 
167). 
 
23. Loranthus ghikae Volkens & Schweinf. in Schweinfurth & Volkens, 
Liste Pl. Somalis 2 (1897). 
– Somalia, Bouka, Kaldesch, 12. 1895, N. & D. Ghika s.n. (holotype, fide 
Polhill in sched.). 
#= Oncocalyx ghikae (Volkens & Schweinf.) M. G. Gilbert (fide Gilbert, 
Polhill & Wiens 1985: 222). 
 
24. Loranthus globiferus A. Rich. var. angustifolius Schweinf. in Bull. 
Herb. Boissier 4, Append. 2: 150 (1896). 
– Ethiopia, Gageros, [1854], Schimper 130 (type). 
= Tapinanthus globiferus (A. Rich.) Tiegh. (fide Polhill in sched.). 
This type specimen is mounted together with Schimper 399, 21. 5. 1852, on 
one sheet. It is not possible to associate the collection numbers with the 
plants. 
– Ethiopia, Amba Marres, 7300' ü. M., 22. 9. 1862, Schimper 888 (syntype). 
= Tapinanthus globiferus (A. Rich.) Tiegh. (fide Polhill in sched.). 
– Ethiopia, in der Ebene Begemder und Dembea, Repp Gabriel, 5700-6500' 
ü. M., 17. 10. 1863, Schimper 1491 (syntype, fide Gilbert 1989: 373, 2 
isosyntypes). 
= Tapinanthus globiferus (A. Rich.) Tiegh. (fide Polhill in sched.). 
– Eritrea, Keren, [Aug. 1861], Steudner 184 (syntype, fide Gilbert 1989: 
373). 
= Tapinanthus globiferus (A. Rich.) Tiegh. (fide Polhill in sched.). - In the 
protologue the collection number of the Steudner specimen is not given. 
 
25. Loranthus goetzei Sprague in Thiselton Dyer, Fl. Trop. Afr. 6(1): 365 
(1910). 
– Tanzania, Ussangu, nördl. Kinga-Berge, Tikurugwa-Berg, 2500 m, 2. 6. 
1899, Goetze 989 (type). 
= Agelanthus fuellebornii (Engl.) Polhill & Wiens (fide Polhill & Wiens 
1992: 163). 
 
26. Loranthus guttatus Sprague in Thiselton Dyer, Fl. Trop. Afr. 6(1): 350 
(1910). 
– Zambia, near Bwana Mkubwa, Rogers 8320 (isosyntype, fide Polhill in 
sched). 
= Tapinanthus dependens (Engl.) Danser (fide Polhill in sched.). 
 
27. Loranthus hirsutiflorus Klotzsch in Peters, Naturw. Reise Mossambique 
6, Bot. 1: 178 (1861). 
– Mozambique, Lupatagebirge, 7. 12. 1844, Peters 8 (holotype, isotype). - 
Also (iso-)syntypes of Loranthus dregei Eckl. & Zeyh. f. subcuneifolius 
Engl. 
= Erianthemum dregei (Eckl. & Zeyh.) Tiegh. (fide Polhill in sched.). In the 
protologue the collection number is not given. 
 
28. Loranthus igneus Sprague in Bull. Misc. Inform. Kew 1912: 232 (1912), 
non (Scheff.) Benth. & Hook. f., nom illegit. 
– Tanzania, Cheminda-Bakary Rondo, 12. 6. 1906, Braun in Herb. Amani 
1169 (holotype, fide Polhill in sched.). 
= Agelanthus igneus (Danser) Polhill & Wiens (fide Polhill & Wiens 1992: 
164). 
 
29. Loranthus ikelembensis De Wild. in Bull. Jard. Bot. État 3: 256 (1911). 
– Zaire, Bombimba, 28. 7. 1909, Laurent 1159 (type). 
= Helixanthera mannii (Oliver) Danser (fide Polhill & Wiens 1992: 168). 
 
30. Loranthus incanus Schum. & Thonn. var. sessilis Sprague in Thiselton 
Dyer, Fl. Trop. Afr. 6(1): 293 (1910). 
– Gabon, Sibangefarm im Gebiet von Munda, 15. 4. 1882, Soyaux 386 (2 
isotypes, fide Polhill in litt.). - Also lecto- and isolectotype of Loranthus 
soyauxii Engl. 
= Phragmanthera capitata (Sprengel) Balle (fide Balle 1982: 29). 
 
31. Loranthus lanceolatus P. Beauv. var. corniculatus Sprague in Thiselton 
Dyer, Fl. Trop. Afr. 6(1): 355 (1910). 
– Togo, Banyatera, um 250 m, 20. 3. 1904, Schröder 224 (holotype, fide 
Polhill in litt.). - Also syntype of Loranthus kerstingii Engl. 
= Tapinanthus globiferus (A. Rich.) Tiegh. (fide Polhill & Wiens 1992: 
173). 
 
32. Loranthus lindensis Sprague in Thiselton Dyer, Fl. Trop. Afr. 6(1): 316 
(1910). 
– Tanzania, bei Kwa-Kihingi (Ungoni), 27. 6. 1903, Busse 3005 (type). 
#= Erianthemum lindense (Sprague) Danser (fide Danser 1933: 54). 
 
33. Loranthus lingelsheimii Pax in Bot. Jahrb. Syst. 39: 615 (1907). 
– Ethiopia, West-Schoa, Hara, 2100 m [2500 m], 25. 3. 1905, Rosen 1909 
(holotype, fide Gilbert 1989: 373). 
= Tapinanthus globiferus (A. Rich.) Tiegh. (fide Polhill & Wiens 1992: 
173). 
 
34. Loranthus longifolius Peter in Repert. Spec. Nov. Regni Veg.  Beih. 
40(2): 178, Anh. 14 (1932). 
– Tanzania, S-Pare Gebirge, Urwald am Berge Stenena, 1910-2000 m, 
15./16. 2. 1915, Peter 8994 (holotype, fide Polhill in sched.). 
= Englerina woodfordioides (Schweinf.) Balle (fide Polhill & Wiens 1992: 
167). 
 
35. Loranthus longipes Baker & Sprague in Thiselton Dyer, Fl. Trop. Afr. 
6(1): 341 (1910). 
– Tanzania, Prov. Zanguebar, [Dar-es-Salaam], Kirk s.n. (isotype, fide 
Polhill in sched.). 
#= Agelanthus longipes (Baker & Sprague) Polhill & Wiens (fide Polhill & 
Wiens 1992: 164). 
 
36. Loranthus lugardii N. E. Br. in Bull. Misc. Inform. Kew 1909: 135 
(1909). 
– Botswana, Lake Ngami, Lugard 32 (isosyntype, fide Polhill in litt.). 
#= Agelanthus lugardii (N. E. Br.) Polhill & Wiens (fide Polhill & Wiens 
1992: 164). 
 
37. Loranthus macrosolen Steudner ex A. Rich., Tent. Fl. Abyss. 1: 340 
(1848). 
– Ethiopia, in provincia Schoata, Schimper 529 (3 isosyntypes, fide Polhill 
in sched.). 
#= Phragmanthera macrosolen (Steud. ex A. Rich.) Balle (fide Polhill & 
Wiens 1992: 171). 
 
38. Loranthus menyharthii Engl. & Schinz ex Schinz in Kaiserl. Akad. 
Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 78: 409 (1906). 
– Mozambique, Sambesi-Mittellauf, Boroma, 6. 1891, Menyharth 936 (3 
isotypes). 
#= Actinanthella menyharthii (Engl. & Schinz ex Schinz) Balle (fide Balle 
1954: 1626). 
 
39. Loranthus mweroensis Baker in Bull. Misc. Inform. Kew 1895: 292 
(1895). 
– Zambia, Tanganjika, [Mwero plateau, west of Lake Tanganyika], Carson 
27 (type). 
#= Globimetula mweroensis (Baker) Danser (fide Polhill & Wiens 1992: 
168). 
 
40. Loranthus myrsinifolius Engl. & K. Krause var. humbertii Balle in Bull. 
Jard. Bot. État 17: 244 (1944). 
– Zaire, montagnes a l'ouest du Lac Kivu, forêt près du marais Kansibi, 
2200 m, 2./3. 1929, Humbert 7515 (type). 
= Agelanthus myrsinifolius (Engl. & K. Krause) Polhill & Wiens (fide 
Polhill & Wiens 1992: 164). - Obviously there exists a clerical error on the 
label with regard to the collection number; in the protologue and also in 
Balle (1948: 333) 7615 is given while all other data on the label of the 
specimen mentioned above agree with the protologue. 
 
41. Loranthus ngamicus Sprague in Thiselton Dyer, Fl. Trop. Afr. 6(1): 310 
(1910). 
– Botswana, Ntschokutsa, 900 m, 14. 12. 1906, Seiner ser. II 124 
(isosyntype, fide Polhill in litt.). 
#= Erianthemum ngamicum (Sprague) Danser (fide Wiens & Tölken 1979: 
30). 
 
42. Loranthus nitidulus Sprague in Thiselton Dyer, Fl. Trop. Afr. 6(1): 283 
(1910). 
– Equatorial Guinea, Fernando Po,  Mann 2346 (isotype, fide Polhill in 
sched.). 
= Phragmanthera polycrypta (F. Didr.) Balle (fide Polhill & Wiens 1992: 
171). 
 
43. Loranthus nyasicus Baker & Sprague in Thiselton Dyer, Fl. Trop. Afr. 
6(1): 371 (1910). 
– Malawi, Nyasaland, Buchanan 1090 (fragment of type, fide Polhill in 
sched.). 
= Agelanthus nyasicus (Baker & Sprague) Polhill & Wiens (fide Polhill & 
Wiens 1992: 164). 
 
44. Loranthus nyikensis Sprague in Thiselton Dyer, Fl. Trop. Afr. 6(1): 315 
(1910). 
= Malawi, McClounie 111 (isotype, fide Polhill in litt.). 
#= Erianthemum nyikense (Sprague) Danser (fide Danser 1933: 54). 
 
45. Loranthus oblongifolius A. Rich., Tent. Fl. Abyss. 1: 342 (1848). 
– Ethiopia, Gapdia, 28. 9. 1838, Schimper It.abyss.II 768 (2 isolectotypes, 
fide Polhill in sched.). - Also syntypes of Loranthus dregei Eckl. & Zeyh. f. 
subcuneifolius Engl. 
= Erianthemum dregei (Eckl. & Zeyh.) Tiegh. (fide Gilbert 1989: 366). 
 
46. Loranthus obovatus Peter in Repert. Spec. Nov. Regni Veg. Beih. 40(2): 
168, Anh. 13 (1932). 
– Tanzania, Landschaft Iraku, Malimo, am Seeufer des Bassoda Merka, 
1880 m, 8. 8. 1926, Peter 43772 (type, fide Polhill in sched.). 
= Agelanthus oehleri (Engl.) Polhill & Wiens (fide Polhill in sched.). 
 
47. Loranthus oleaefolius Cham. & Schlechtend. var. luteus Neusser in 
Mitt. Bot. Staatssamml. München 1: 339 (1953). 
– Namibia, Bezirk Karibib, Farm Nudis, auf Bäumen im Swakoptal, 13. 2. 
1953, Walter & Walter 1288 (isosyntype). 
= Tapinanthus oleifolius (J. C. Wendl.) Danser (fide Balle 1968: 182). 
 
48. Loranthus opacus Sprague in Thiselton Dyer, Fl. Trop. Afr. 6(1): 300 
(1910). 
– Gabon, Gaboon River, Mann s.n. (isotype, fide Polhill in sched.). 
= Globimetula dinklagei (Engl.) Danser (fide Polhill & Wiens 1992: 174). 
 
49. Loranthus ophiodes Sprague in Bull. Soc. Bot. France 58: 202 (1912). 
– Burkina Faso, Gourma, [environs de Diapaga], [12.-18. 7. 1910], 
Chevalier 24438 (isosyntype). 
#= Tapinanthus ophiodes (Sprague) Danser (fide Polhill & Wiens 1992: 
174) 
 
50. Loranthus oreophilus Oliv. in J. Linn. Soc., Bot. 7: 196 (1864). 
– Cameroon, Cameroon mountains, Mann 2163 (isotype, fide Polhill in 
sched.). 
#= Globimetula oreophila (Oliv.) Danser (fide Danser 1933: 55). 
 
51. Loranthus pachycladus Sprague in Bull. Misc. Inform. Kew 1912: 233 
(1912). 
– Tanzania, Ussangu, nördl. Kinga-Berge, Tyuni-Berg, 5. 6. 1899, Goetze 
1003 (holotype, 2 isotypes, fide Polhill in litt.). 
#= Globimetula pachyclada (Sprague) Danser (fide Danser 1933: 55). 
 
52. Loranthus pennatulus Sprague in Thiselton Dyer, Fl. Trop. Afr. 6(1): 
324 (1910). 
– Kenia, 1902, Elliott 23 (syntype). 
#= Agelanthus pennatulus (Sprague) Polhill & Wiens (fide Polhill & Wiens 
1992: 164). 
 
53. Loranthus pentagonia DC., Prodr. 4: 303 (1830). 
– Senegal, "Herb. Kunth. Senegal. En Hudes Paris 1821" (isosyntype?, fide 
Polhill in sched.). 
#= Tapinanthus pentagonia (DC.) Tiegh. (fide Polhill in sched.). 
It is uncertain whether the specimen in B is the type specimen collected by 
Perrottet & Leprieur. There are six specimens of L. pentagonia on the 
microfiche edition of the Candolle herbarium at G, four without any 
collection data, one specimen labeled "Perrottet 8. Janvier 1825 Nr. 163" 
and another "Perrottet & Leprieur 1829". None is marked as type specimen. 
 
54. Loranthus pentheri Schlechter in J. Bot. 36: 376 (1898). 
– Zimbabwe, Ligombwe, [in arboribus prope Ligombwe, in terra 
Matabeleland], 26. 6. 1895, [A. Penther] (holotype, fide Polhill in sched.). 
= Plicosepalus kalachariensis (Schinz) Danser (fide Polhill in sched.). 
 
55. Loranthus platyphyllus Hochst. ex A. Rich., Tent. Fl. Abyss. 1: 341 
(1848). 
– Ethiopia, [prope Tchélatchékanné, in valle fluvii Taccazé], 19. 4. 1841, 
Schimper 1584 (2 isotypes; holotype in P, fide M. G. Gilbert 1989: 372). 
#= Agelanthus platyphyllus (Hochst. ex A. Rich.) Balle (fide Balle 1970: 2). 
 
56. Loranthus poecilobotrys Werth, Veg. Sansibar: 35 (1901). 
– Tanzania, Zanzibar, Insel Sbr. Mitschawi (Sandland), Werth s.n. 
(holotype). 
= Oncella ambigua (Engl.) Tiegh. (fide Polhill & Wiens 1992: 169). 
 
57. Loranthus prunifolius E. Mey. ex Harv. in Harvey & Sonders, Fl. Cap. 
2: 578 (1862). 
– South Africa, Z.n.N. n.E. [C. Zeyher] (2 isosyntypes). 
#= Agelanthus prunifolius (E. Mey. ex Harv.) Polhill & Wiens (fide Polhill 
in sched.). 
It is uncertain, if these 2 collections are type specimens. On the label of one 
of them only the abbreviation "Z.n.N. n.E." is readable, which stands for 
"Zeyher's neuere Nachlieferung von Pflanzen, welche nicht mehr in E. Z. 
Enumeratio aufgenommen wurden, aber in den von Drège gemachten 
Vertheilungen der E.Z.schen Pflanzen vorkommen..." (fide Drège 1847: 
599). On the label of the other specimen there are no data besides the 
species name. The handwriting is probably the same as illustrated and 
explained to belong to an amanuensis byGunn & Codd (1981: 145, 146). 
Possibly C. Zeyher alone is the collector of these specimens while in 
Harvey (1862) Drège, Eckl. & Zeyh. and H. Hutton are given as collectors. 
The difficulty to determine the collector(s) (Drège, Eckl. & Zeyh. or only 
Zeyh.) is discussed by Gunn & Codd (1981). 
 
58. Loranthus quinquangulus Engl. & Schinz ex Schinz in Kaiserl. Akad. 
Wiss. Wien, Math.-Naturwiss.Kl., Denkschr. 78: 409 (1906). 
– Mozambique, Sambesi-Mittellauf, Boruma, Nhaondua, 5. 1892, 
Menyharth 615 (isotype, fide Polhill in sched.). 
= Tapinanthus oleifolius (J. C. Wendl.) Danser (fide Polhill & Wiens 1992: 
174). 
 
59. Loranthus ramulosus Sprague in Bull. Misc. Inform. Kew 1911: 182 
(1911). 
– Kenya, [Mombasa-Lamu, Whyte] (isosyntype). 
#= Englerina ramulosa (Sprague) Polhill & Wiens (fide Polhill & Wiens 
1992: 167). 
 
60. Loranthus regularis Steud. ex Sprague in Thiselton Dyer, Fl. Trop. Afr. 
6(1): 287 (1910). 
– Ethiopia, am Tana-See bei Gorgora, Steudner 173 (type). 
#= Phragmanthera regularis (Steud. ex Sprague) M. G. Gilbert (fide Gilbert, 
Polhill & Wiens 1985: 224). 
 
61. Loranthus remotus Baker & Sprague in Thiselton Dyer, Fl. Trop. Afr. 
6(1): 327 (1910). 
– Mozambique, Shupanga, 8. 1858 [this date is written on a drawing on the 
sheet], Kirk 40 (isotype). 
#= Vanwykia remota (Baker & Sprague) Wiens (fide Wiens & Tölken 
1979: 29). 
 
62. Loranthus roseus Klotzsch in Peters, Naturw. Reise Mossambique 6, 
Bot. 1: 177 (1861). 
– Mozambique, Lourenzo-Marques, (Delagoa Bay), 11. 1846, Peters s.n. 
(holotype). - Also syntype of Loranthus dregei Eckl. & Zeyh. f. obtusifolius 
Engl. 
= Erianthemum dregei (Eckl. & Zeyh.) Tiegh. (fide Polhill in sched.). 
 
63. Loranthus rubroviridis Oliver var. bechuanicus Sprague in Thiselton 
Dyer, Fl. Trop. Afr. 6(1): 299 (1910). 
– Botswana, [Bechuana country, Eastern Bamanguato Territory; between 
Nagatatollo and Henrys-pan], Holub s.n. (isotype). 
=  rubroviridis Tiegh. (fide Polhill in litt.). 
 
64. Loranthus sambesiacus Engl. & Schinz ex Schinz in Kaiserl. Akad. 
Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 78: 409 (1906). 
– Mozambique, Sambesigebiet, am Mittellauf, Boruma, Nhaondua, 6. 1891, 
Menyharth 939 (isotype). 
#= Agelanthus sambesiacus (Engl. & Schinz ex Schinz) Polhill & Wiens 
(fide Polhill & Wiens 1992: 165. 
 
65. Loranthus schimperi Hochst. ex A. Rich., Tent. Fl. Abyss. 1: 341 
(1848). 
– Ethiopia, [circa Adoua et in valle rivi Chahagni], 31. 12. 1837, Schimper 
158 (3 isosyntypes, fide Polhill in sched.). 
#= Oncocalyx schimperi (Hochst. ex A. Rich.) M. G. Gilbert. (fide Gilbert, 
Polhill & Wiens 1985: 222). - There is probably another type specimen 
without collection number on the label kept in B. 
 
66. Loranthus stuhlmannii Engl. var. somalensis Engl. ex Sprague in 
Thiselton Dyer, Fl. Trop. Afr. 6, 1: 331 (1910). 
– Ethiopia, Webi [to] Abdallah, Keller 218 (isotype). 
= Oncocalyx fischeri (Engl.) M. G. Gilbert (fide Gilbert 1989: 369). 
 
67. Loranthus sublilacinus Sprague in Thiselton Dyer, Fl. Trop. Afr. 6(1): 
275 (1910). 
– Cameroon, Johann-Albrechtshöhe, 12. 1896, Staudt 757 (syntype, 
isosyntype, fide Polhill in litt.). - Also syntypes of Loranthus rosaceus Engl. 
= Helixanthera mannii (Oliv.) Danser (fide Polhill & Wiens 1992: 168). 
– Cameroon, Bipindi [Bipinde], 3. 1902, Zenker 2518 (syntype, fide Polhill 
in sched., isosyntype). 
= Helixanthera mannii (Oliv.) Danser (fide Polhill & Wiens 1992: 168). 
 
68. Loranthus talbotiorum Sprague in Thiselton Dyer, Fl. Trop. Afr. 6(1): 
1026 (1913). 
– Nigeria, [Oban district], Talbot & [Talbot] 1281 (isotype, fide Polhill in 
sched.). 
#= Phragmanthera talbotiorum (Sprague) Balle (fide Balle 1956: 168). 
 
69. Loranthus triplinervius Baker & Sprague in Thiselton Dyer, Fl. Trop. 
Afr. 6(1): 385 (1911). 
– Tanzania, Versuchsgarten in Dar-es-Salaam, 20. 2. 1902, Holtz 553 
(type). 
#= Englerina triplinervia (Baker & Sprague) Polhill & Wiens (fide Polhill 
& Wiens 1992: 167). 
– Tanzania, Dar-es-Salaam, 31. 3. 1894, Stuhlmann 7614 (type). 
#= Englerina triplinervia (Baker & Sprague) Polhill & Wiens (fide Polhill 
& Wiens 1992: 167). 
– Tanzania, Dar-es-Salaam, 6. 4. 1894, Stuhlmann 7666 (type). 
#= Englerina triplinervia (Baker & Sprague) Polhill & Wiens (fide Polhill 
& Wiens 1992: 167). 
– Tanzania, 1894, Stuhlmann 7667 (type). 
#= Englerina triplinervia (Baker & Sprague) Polhill & Wiens (fide Polhill 
& Wiens 1992: 167). 
– Tanzania, Dar-es-Salaam, 7. 4. 1894, Stuhlmann 7673 (type). 
#= Englerina triplinervia (Baker & Sprague) Polhill & Wiens (fide Polhill 
& Wiens 1992: 167). 
– Tanzania, Dar-es-Salaam, 4. 1894, Stuhlmann 7790 (type). 
#= Englerina triplinervia (Baker & Sprague) Polhill & Wiens.  
70. Loranthus undulatus E. Mey. ex Harv. in Harvey & Sonders, Fl. Cap. 2: 
577 (1862). 
– South Africa, [between Natvoet and the Gariep, and near Verleptpram], 
Drège P. b. Sp. [written only on the label of one specimen] (2 isosyntypes?, 
fide Polhill in sched.). 
#= Plicosepalus undulatus (E. Mey. ex Harvey) Tiegh. (fide Polhill in 
sched.). 
It is uncertain whether these specimens are type specimens since the name 
of the species is missing on both labels and "Drège P. b. Sp." on one label 
was probably written by an amanuensis (cf. Gunn & Codd 1981: 145, 146 
and sub no. 57, see above); there are no collection data on the labels. 
 
71. Loranthus viridizonatus Werth, Veg. Sansibar 87 (1901). 
– Tanzania, [Zanzibar], Werth s.n. (holotype). 
= Agelanthus sansibarensis (Engl.) Polhill & Wiens (fide Polhill & Wiens 
1992: 165). 
 
72. Loranthus woodfordioides Schweinf. in Höhnel, Rudolph-Stephanie-See 
856 (1892). 
– Kenya, Guasso Narok, 2000 m, 1887/1888, Höhnel 39 (holotype, fide 
Polhill in sched.). 
#= Englerina woodfordioides (Schweinf.) Balle (fide Balle 1958: 108). 
 
73. Phragmanthera rufescens (DC.) Balle subsp. longiflora Balle in Satabié 
& Leroy, Fl. Cameroun 23: 39 (1982). 
– Cameroon, [Strasse Kribi - Jaunde nördlich des Njong ca. 40 km SSW 
Jaunde.], [14. 1. 1914], Mildbraed 7763 (holotype, fide Balle 1982: 39). 
#= Phragmanthera longiflora (Balle) Polhill & Wiens (fide Polhill & Wiens 
1992: 171). 
 
74. Plicosepalus robustus Wiens & Polhill in Nordic J. Bot. 5: 221 (1985). 
– Ethiopia, Sidamo Region, east foot of Amaro Mts. Keli, 24. 1. 1953, 
Gillett 14869 (type). 
 
75. Tapinostemma meridianum Danser in Notizbl. Bot. Gart. Berlin-
Dahlem 11: 216 (1931). 
– Tanzania, Kilimadinde, Südufer des Msombe-Flusses zwischen Irangali 
und Uvimbi, 27. 5. 1905, Prittwitz 175 (holotype). 
#= >Plicosepalus meridianus (Danser) Wiens & Polhill (fide Gilbert, Polhill 
& Wiens 1985: 222). 
– Tanzania, Dornbusch bei Kirungule (Ugogo), 14. 8. 1900, Busse 233 
(paratype). 
#= Plicosepalus meridianus (Danser) Wiens & Polhill (fide Gilbert, Polhill 
& Wiens 1985: 222). 
– Tanzania, Fischer 535 (paratype). 
#= Plicosepalus meridianus (Danser) Wiens & Polhill (fide Gilbert, Polhill 
& Wiens 1985: 222). 
– Tanzania, 7. 3. 1906, Jaeger 38 (paratype). 
#= Plicosepalus meridianus (Danser) Wiens & Polhill (fide Gilbert, Polhill 
& Wiens 1985: 222). 
 
Acknowledgements 
The author is grateful to Paul Hiepko for help in taxonomic questions, for 
correcting and improving the English text and support in many other ways 
throughout this work. It is a pleasure for me to thank Roger Polhill, Kew, 
for many important comments. 
 
Index to the type collections: 
Beccari 82 (15). 
Braun in Herb. Amani 1169 (28), 1970 (19). 
Buchanan 1090 (43). 
Bunting 165 (7). 
Busse 233 (75), 764 (17), 2949 (8), 3005 (32). 
Carson 27 (39), s.n. (9). 
Cecil 96 (10). 
Chevalier 24438 (49). 
Demeuse 119 (5). 
Drège s.n. (70). 
Drummond & Hemsley 2577 (79). 
Ecklon & C. Zeyher 2284 (13). 
Ellenbeck 797 (14), 1303 (14). 
Elliott 23 (52). 
Engler 707 (19). 
Fischer 535 (75). 
Ghika, N. & D. s.n. (23). 
Gillett 14869 (74). 
Goetze 162 (18), 989 (25), 1003 (51), 1014 (18). 
Gossweiler 149 (6), 473 (6). 
Hildebrandt 517 (15). 
Holst 2946 (3). 
Holtz 553 (69). 
Holub s.n. (63). 
Humbert 7515 (40), 14060 (1). 
Jaeger 38 (75). 
Keller 218 (66). 
Kirk 40 (61), s.n. (12), s.n. (35). 
Laurent 1159 (29). 
Lugard 32 (36). 
Mann 2163 (50), 2346 (42), s.n. (48). 
McClounie 111 (44). 
Menyharth 615 (58), 936 (38), 939 (64). 
Mildbraed 7763 (73). 
Penther s.n. (54). 
Peter 1888 (22), 8994 (34), 43772 (46), 44752 (4). 
Peters 8 (27), s.n. (62). 
Polhill & Paulo 1694 (80). 
Prittwitz 175 (75). 
Rogers 8318 (2), 8320 (26). 
Rosen 1909 (33). 
Schimper 130 (24), 158 (65), 529 (37), 768 (45), 888 (24), 1491 (24), 1584 
(55). 
Schinz 291 (20), 294 (11). 
Schlieben 3285 (76), 4012 (77), 6121 (78). 
Schröder 224 (31). 
Schweinfurth 2193 (15). 
Seiner ser. II 124 (41). 
Soyaux 386 (30). 
Staudt 757 (67). 
Steudner 173 (60), 184 (24). 
Stuhlmann 7614 (69), 7666 (69), 7667 (69), 7673 (69), 7790 (69). 
Talbot & Talbot 1281 (68). 
Volkens 1934 (16). 
Walter & Walter 1288 (47). 
Welwitsch 4849 (21). 
Werth s.n. (56), s.n. (71). 
Whyte s.n. (59). 
Zenker 2518 (67). 
Z.n.N.n.E. [Zeyher] (57). 
Herb. Kunth. Senegal. En Hudes Paris (53). 
 
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Address of the author: 
R. Gebauer, c/o Botanischer Garten und Botanisches Museum Berlin-
Dahlem, Königin-Luise-Straße 6-8, D-14195 Berlin.