Electronic supplement to: 
King-Jones (née Hunger), S.: Revision of Pluchea Cass. (Compositae, Plucheeae) 
in the Old World. – Englera 23: 1-136. 2001.


Specimens examined


1. Pluchea arabica (Boiss.) Qaiser & Lack

OMAN: 40 km W of Salalah., 2 km from Mugsail, 16°45'N, 53°40'E, 9.11.1984, I. McLeish 
322 (E); Dhofar: Disected plain NE of Mirbat [Mirbat: 16°58'N, 54°43'E], 2.8.1985, A. G. 
Miller 7231 (E); Dhofar: Sea cliffs nr Ras Hamar, W of Raysut [c. 16°56'N, 53°50'E], 
28.9.1979, A. G. Miller 2314 (E, K); W Hinua E of Taqah [17°00'N, 54°25'E], 30.5.1985, A. 
G. Miller 7037 A (K); Dhofor: Mirbat, Wadi 5 km ENE of Mirbat, 17°01'N, 54°44'E, 
19.9.1984, A. G. Miller 6181 (E); Jabal Qinguri, 17°02'N, 55°01'E, 4.11.1983, R. M. Lawton 
2521 (BM 21044); Wadi on north side of Jebel Qara, [c. 17°10'N, 54°30'E], 9.3.1979, R. 
Whitecombe 506 (E); 9.3.1979, R. Whitecombe 514 (E); 72 km of Ashinhaib to Zeak, 
17°20'N, 54°10'E, 27.4.1985, McLeish 520 (E); Wadi Dawkah, N foothills of Jabel Qara, 
17°23'N, 54°00'E, 21.9.1988, M. D. Gallagher 6898/7 (E); Jebel Qara, 26.10.1977, R. M. 
Lawton 1857 (BM 21043, K); 7.10.1943, D. Vesey-Fitzgerald 1254/4 (BM 21153); Dhofar: 
Plateau above Schuwaymiyah [17°53'N, 55°54'E], 18.9.1989, A. G. Miller & J. A. Nyberg 
9426 (E); Dhofar: Ayoon cross roads [Ayun: 17°15'N, 53°53'E], 23.9.1978, A. Berkeley 109 
(E); 6.5.1978, A. Berkley 15 (BM 20926); Wadi 18 km S of Thamrait on Salalah road [c. 
17°30'N, 54°00'E], 15.9.1984, A. G. Miller 6044 (K); Sawqirah [18°07'N, 56°32'E], 
29.9.1984, A. G. Miller 6475 (K); Dhofar, Assubaithi, 1992, M. Jongblood (K); Jiddat al 
Harasis: Huqf Escarpment, 10 km S of Myhoos [c. 19°50'N, 56°30'E], 22.9.1989, A. G. Miller 
& Nyberg 9501 (K); Yalooni, SE of Haima [19°56'N, 56°19'E], 28.4.1982, J. R. Maconochie 
3424 (E); Nr. Haql, Masirah Is., 20°22'N, 58°47'E, 4.2.1982, M. D. Gallagher 6356/2 (E); 25 
km N Sur on Al Kamil Rd [22°20'N, 59°30'E], 2.4.1982, J. R. Maconochie 3307 (E); Adam to 
Nizwa, 30 km before Nizwa [c. 22°30'N, 57°30'E], 15.10.1984, A. G. Miller 6583 (K); W 
Rafsah, SW of Sur [c. 22°50'N, 59°20'E], 16.3.1976, A. Radcliffe-Smith 3923 (K); Amlah near 
Ibri, Wadi al Ayn [Ibri: 23°15'N, 56°16'E], 12.1.1978, R. Whitecombe 67 (E); Muscat: 10 km 
W of Muscar, [23°37'N, 58°38'E], 18.3.1975, T. G. Rubens 81 (E); Lansab industrial area, 
23°32'N, 58°18'E, 5.6.1987, M. D. Gallagher 7976/16 (E); E Batinah, 5 miles SE of Seeb 
Intnl. Airport [c. 23°45'N, 58°20'E], 26.2.1976, A. Radcliffe-Smith 3635 (E, K); Nizwa Agric 
Institut farm, Firq, 23°47'N, 57°47'E, 17.10.1981, J. R. Maconochie 2862 (BM 21057, E); 
Wadi Jizzi, near the frontier check-post, 50 km E of Buraimi City, highway to Sohar 
[24°23'N, 56°35'E], 10.11.1986, L. Boulos & A. El-Ghonemy 16318 (BM 20932).

YEMEN: Hadramaut, 25 km NE of Ghayl Ba Wazir, steep, rocky wadi on the W side of and 
tributary to Wadi Arf, 4 km N of the oasis and entrance into the gorge, 14°59'N, 49°30'E, 
21.9.1998, N. Kilian & P. Hein NK 5027 (B); Mahrah: 16 km from Sayhut along road to 
Qishn, 15°15'N, 51°21'E, 16.10.1992, Thulin, Eriksson, Gifri & Langstroem 8428 A (UPS); 
Nitaura bei Ras Fartak [c. 15°38'N, 52°13'E], 7.-8.3.1899, St. Paulay (HBG, WU); Al Mahra, 
mountain range W of Qishn, 15°22'N, 51°36'E., 24.9.1998, N. Kilian, P. Hein & S. Ghazanfar 
NK 5091 (B); Al Mahra, Jabel Sharwayn 10 km W of Qishn, 15°24'N, 51°35'E, 28.9.1998, P. 
Hein 5098 (herb. P. Hein); Gischin, Hadramaut [?Qishn: 15°25'N, 51°40'E], 1902, W. Hein 
(WU); Al Mahra, wadi just S of Nishtun, 15°48'N, 52°11'E, 27.9.1998, N. Kilian, P. Hein & 
S. Ghazanfar NK 5239 (B); Al Mahra, track between Ghaydah to Sarif Spring, lower Wadi 
Hablah 40 km NNW of Ghaydah, 16°35'N, 52°03'E, 25.9.1998, P. Hein 4941 (herb. P. Hein); 
Al Mahra, track from Ghaydah to Sarif spring, lower Wadi Hablah, 16°40'N, 52°E, 25.9.1998, 
N. Kilian, P. Hein & S. Ghazanfar NK 5143 (B); Wadi Gabur, Umgebung von Gischin, 
Hadramaut, 7.2.1902, W. Hein (WU); Gatgit, Gischin, Hadramaut, 27.3.1902, W. Hein (HBG, 
WU).


2. Pluchea arguta Boiss.

PAKISTAN: Cape Monze W Karachi [24°49'N, 66°40'E], 6.4.1965, K. H. Rechinger 27526 (B, 
G, M); 4.4.1965, D. Podlech 10073 (M), 6.4.1965, J. Lamond 241, 247 (C, E); Karachi Distr., 
W. Koelz 7568 (G, S); 8 miles from Paradise point on way to Cape Monze, Karachi, 
15.6.1969, Sultan-ul-Abedin 3428 (B); Karachi to Dadu [Karachi: 24°51'N, 67°02'E], 
4.5.1965, J. Lamond 813 (E), 9.4.56, S. M. H. Jafri [?] 1316 (E); 1 mile from Hub Choki, on 
way to Sonmiani [25°05'N, 66°55'E], 24.2.1969, S. A. Lavogue [?] & Sultan ul Abedin 1180 
(E); E Thano Bula Khan, 25°23'N, 67°51'E, versus Kotri 25°22'N 68°28'E, 4.5.1965, 
Rechinger 28661 (B, G, K, M, S); Ganzar hills in gorges near Hoshab on way to Padarak 
[26°01'N, 63°56'E], 23.9.1986, A. Ghafoor & S. Omeh 1750.A (G); A. Ghafoor & S. Omeh 
1765.A (G); c. 3 km from village Kikin, bet. village Shahrak + Turbat [26°02'N, 63°20'E], 
13.3.1990, A. Ghafoor & S. M. Goodmann 4595 (E); mt. Salinorum, Sakesar [32°27'N, 
72°02'E], 14.9.1902, J. R. Drummond 14569 (E, K); between Bannu and Kohat [Kohat: 
33°35'N, 71°26'E], 3.2.1934, R. N. Parker 3301 (A, K); Punjab, Distr. Mianwali, near 
Kalabagh [32°58'N, 71°34'E], 29.12.1, J. R. Drummond 13788 (E).

INDIA: Bombay [18°56'N, 72°51'E], N. A. Dalzell (K); Thimper Lake, Kinghar Lake [24°54'N, 
68°03'E], Makli hills [26°39'N, 80°28'E], 24.2.1896, Herb. Brandis 3 (HBG); N W India, 
1871, J. L. Stewart (E); Punjab, J. R. Drummond 25756 (G, K).


3. Pluchea baccharoides (F. Muell.) F. Muell ex Benth.

AUSTRALIA: Waverley Downs to Hungerford [29°06'S, 143°54'E], 10.1912, J. L. Boorman (B, 
BM[labeled b] 21071); Paroo River district [30°05'S, 144°05'E-29°34'S, 144°17'E], 10.1912, 
J. L. Boorman (G).
QUEENSLAND: 4.5 km E of Moray Downs HS, 21°57'S, 146°40'E, 5.4.1992, E. J. Thompson & 
B. K. Simon BUC 346 (MEL 1616644); 15 miles ENE of “Surbiton” Stn., S Kennedy Dist. 
[23°09'S, 146°37'E], 25.9.1964, L. G. Adams 1318 (CANB 152193).
WESTERN AUSTRALIA: Coolgardie Goldfields [30°57'S, 121°09'E], 10.1909, Maiden (C); 
Esperance [33°49'S, 121°52'E], 11.1909, Maiden (C).


4. Pluchea bojeri (DC.) Humbert

ETHIOPIA: Gondaraba [4°58'N, 36°49'E], 1939, Corradi 1949 (W); Kibish-Wells, [5°14'N, 
35°50'E], 12.6.1953, S. H. Padwa 270 (B, BR).

CONGO: Zone d’Ubundu, 0°8'N, 25°4'E, 20.11.1982, J. Lejoy 82/646 (BR); Katenga [5°01'S, 
28°49'E], 9.4.1926, W. Robyns 1861 (BR); Malele [5°22'S, 19°22'E], 7.1.1919, F. Vermoesen 
1343 (BM 21152, BR); Matadi [5°50'S, 13°32'E], 6.10.1930, Vanderyst 25880, 25881, 25978, 
25989 (BR), 1913, Verscheuren 873 (BR), 1915, Bequert 7805 (BR); Boma [5°51'S, 
13°03'E], 3.12.1918, F. Vermoesen 1048 (BR), 10.12.1918, F. Vermoesen 1120 (BR), 28. 
6.1919, F. Vermoesen 2500 (BR), 10.1924, Vanderyst 13908 (BR), 31.06.1913, Bequert 525 
(BR); Babana [6°01'S, 12°24’E], 1919, Vermoesen 2546 (G), 27.6.1895, A. Dewevre 52 (BR); 
Congo, 1816, C. Schmith (BM 21126); Gue de la Dikuluwen, route de Kayombo, 10°16'S, 
26°33'E, 23.7.1952, A. Schmitz 4020 (BR); Mwashya [10°42'S, 27°20'E], 26.2.1972, Thoen T 
5384 (BR); Parc Nacional de L’Umpemba, 16.6.1948, de Witte 3970 (BR).

KENYA: Oloregasailie [c. 1°35'S, 36°26'E], 29.8.1943, Bally 2699 (G); Ft. Shomboli Ngara 
Nyiro [2°08'S, 36°05'E], 24.9.1944, P. R. Bally 3806 (G); Galana Ranch [c. 2°40'S, 39°30'E], 
9.8.1975, Bally 16823 (BR); Tsavo Royal National Park [c. 2°45'S, 39°00'E], 10.1964, R. 
Williams & D. Sheldrick 99 (BR), 30.9.1965, H. P. Hucks 442 (A); Kilimandscharo [c. 
3°04'S, 37°22'E], 4.1894, G. Volkens 2218 (G); Jilore Mida, SW of Malindi [3°12'S, 39°54'E], 
18.3.1973, G. W. Sangai EA 15659 (M); Mombasa [4°04'S, 39°40'E], 1847-52, M. Boivin (P 
11561); Marafa, 25 ml NW of Malindi [3°02'S, 39°58'E], 21.11.61, R. Polhill & S. Paulo 830 
(B, BR).

TANZANIA: Serenela River, Banagi area [c. 2°16'S, 34°51'E], 3.11.1953, R. Tanner 1663 
(BR); Pare: bei Makania [4°20'S, 37°51'E], 5.2.1915, A. Peter 40 (BR); Pare, Kissangara, 
4.2.1915, A. Peter 210 (BR); Gombelo, [4°41'S, 38°37'E], 2. 1893, C. Holst 2171 (HBG, M); 
Pemba Island [5°08'S, 39°40'E], 1928, Vaughan 290 (BM 21151); Kwernuyu in wet valley, 
[5°10'S, 38°48'E], 8.3.1957, R. E. S. Tanner 3480 (B, BR); Madanga [5°21'S, 38°59'E], 
4.4.1957, R. E. S. Tanner 3470 (B, BR), 17.12.1957, R. E. S. Tanner 3905 (B, BR); Mkwaja, 
Sima [5°47'S, 38°51'E], 13.6.1957, R. E. S. Tanner 3570 (B, BR); Sansibar [c. 6°10'S, 
39°20'E], 16.11.1888, Stuhlmann (HBG), 1873, Hildebrandt 1018 (BM 21005), 2.10.1963, H. 
Faulkner 3285 (BR); Mkata, Uferwald des Mkata Flusses, [6°45'S, 37°21'E], 23.11.1925, A. 
Peter 32389 (B); Dar es Salam, [6°48'S, 39°15'E], 3.4.1968, B. J. Harris 1520 (C); Univ. 
Campus, Dar es Salaam, 9.10.1978, Cecilia Markström MN 89 (LD); about 20 miles W of 
Dar es Salaam [c. 6°48'S, 39°20'E], 11.8.1971, O. J. Hansen 401 (C); Kundushi, 24.2.1988, 
Breyne 5528 (BR); Uzaramo, Musua, Bagalla km 99-99.5 südl. d. Eisenbahn, [6°48'S, 
38°34'E], 6.11.1925, A. Peter 31879 (B); Uluguru Gebirge, Morogoro [6°49'S, 37°40'E], H. J. 
Schlieben 3363 (B, BR, G); Mwane about 40 mls from Morogoro, 11.2.1970, Borota 69 (BR); 
Karema [6°49'S, 30°26'E], 1891, R. P. Sacleux (P 11571); Ruaha River on the road to 
Dodoma 18.6.1938, Pole Evans & J. Erens 804 (BR, P 11567); Morogoro: Uluguru-Gebirge, 
Nordwestseite, [c. 7°10'S, 37°40'E], 1.2.1932, H. J. Schlieben 3363 (B, BM 21111, BR, M); 
Kideté, 6.6.1921, Swynnerton 820 (BM 21131); Tumba, [7°31'S, 31°40'E], 9.2.1950, A. A. 
Bullock 2400 (B, BR), 13.3.1951, A. A. Bullock 3774 (B, BR); Bezirk Mahenge-Kilwa, 
Gegend des Zusammenflusses von Kilombero u. Luwegu zum Rufiji, [c. 8°31'S, 37°22'E], 
14.6.1932, H. J. Schlieben 2398 (B, BR); Tanga Prov. Pangani, 5.11.1956, R. E. S. Tanner 
3339 (B, BR).

ANGOLA: Ambiz [7°51'S, 13°07'E], 11.1853, Welwitsch 3927 (BM 21166, G, P 11554); 
Luanda, Calumbo [9°09'S, 13°25'E], Welwitsch 3924 (BM 21163, BR, G, P 11551); Loanda 
[11°02'S, 14°54'E], 1903, Gossweiler 257 (P 11549), 1935, Gossweiler 10472 (BM 20988); 
Lobito Bay [12°20'S, 13°34'E], 8.3.1927, H. Lynes (BM 21055); Benguela [12°34'S, 
13°24'E], 1857-8, Wawra 246 (W); zwischen Benguela u. Lobito, 29.11.1951, H. Hess 51/5 
(M); Mossamedes [15°11'S, 12°09'E], 1859, Welwitsch 3922 (BM 21162, G, P 11550).

MALAWI: Lilongwe [13°58'S, 33°49'E], 5.5.1984, Patel, Seyani & Banda 1502 (C); Liwonde 
at the Hot Springs, [15°01'S, 35°15'E], 27.3.1984, Banda & Kaunda 2126 (C).

MOZAMBIQUE: Negomano, 11°20'S, 38°30'E, 1.11.1960, A. Gomes & Sousa 4593 (M); 
Chemba [17°09'S, 34°53'E], 1929, P. Lesne (P 11576); Gaza: Canicade [c. 24°56'S, 33°42'E], 
29.7.1969, M. F. Correia & A. Marques 965 (M); Namaacha [25°58'S, 32°01'E], 4.6.1974, A. 
Marques 2464 (M); Maputo [25°58'S, 32°35'E], 7.4.1947, R. M. Hornby 2678 (P 11575).

ZAMBIA: Mt Makulu Res. Stn. 12 mls. S. Lusaka [c. 15°01'S, 28°14'E], 9.6.1956, A. Angus 
1335 (BM 20918, BR); Gwembe Valley [16°30'S, 28°00'E], 31.3.1952, F. White 2377 (BR); 
above R. Zambezi at Katombora [17°50'S, 25°24'], 6.1.1953, A. Angus 1106 (BM 20917, 
BR); Manora?, 2.3.1958, R. Goodier & J. B. Phipps 243 (BR).

ZIMBABWE: Kariba [16°31'S, 28°48'E], 4.1960, B. Goldsmith 52/60 (LD, M); Nyanyanya 
River, near Kariba, 2.2.1958, R. B. Drummond 5440 (BR); Wankie [18°22'S, 26°29'E], 
21.6.1934, Eyles 8075 (BM 20980); Zambesie, G. le Testu 710 (BM 21143, P 11577); 
Chitsas, 14.6.1950, N. C. Chase 2346 (BM 20952); Lower Sabi, Devuli river, 1.2.1948, H. 
Wild 2459 (BR); Khami, 25.4.1946, H. Wild 1058 (BR).

NAMIBIA: Ososouu River at Otjiwero, [17°57'S, 13°22'E], 3.4.1957, de Winter & Leistner 
5363 (B, M); Kaokoveld: Otju am Rivier [18°15'S, 13°15'E], 28.1.1958, H. Merxmüller 1424 
(M); Kaokoveld, Otjihu [18°20'S, 13°20'E], 2.5.1957, de Winter & Leistner 5657 (B, M); am 
Ufer des Kunene bei Otjinungua, 12.6.1965, W. Giess 8937 (M); Sarusas Ost [c. 18°50'S, 
12°30'E], 26.4.1966, W. Giess 9414 (M); Otjitambi am Hondotorivier [19°49'S, 15°10'E], 
17.6.1960, W. Giess 3147 (M).

BOTSWANA: Nata River, 20°15'S, 26°11'E, 9.4.1983, P. A. Smith 4215 (M).
S Africa: Koningsmark, Dongola [22°16'S, 29°34'E], 7.1.1948, J. B. Pole Evans 4541 (C, M); 
Dongola Reserve, 3.12.1942, R. A. Dyer 4303 (M); Chipise [22°36'S, 30°10'E], 7.2.1959, 
Meeuse 10597 (B, BR, M); Mkuze Bridge on Mkuze – Candover Rd., 23.1.1963, D. Edwards 
2890 (M); Umhlatuzi River area near U. C. Z., 15.9.1966, H. J. T. Venter 2697 (M).

MADAGASCAR: baie de Diégo-Suarez [Antseranan: 12°19'S, 49°17'E], 1848, Boivin 2296 (P); 
Vohémar [13°22'S, 50°00'E], 1837, Richard 7 (C), Richard (G), Richard 558 (P), 109 (P), 36 
(P); 1847-1852, Boivin 2395 (P); Maromandia [14°10'S, 48°06'E], 31.3.1923, Decary 1634 
(L, P); Manambaro [14°25'S, 48°13'E], 9.-18.6.1968, D. Seligson 679 (A, P); Tsingy de 
Namoroka [16°28'S, 45°20'E], Service Forestier de Madagascar 7-RN (P); Vallee de la 
Betsiboka  Ambato-Boeni [16°30'S, 46°38'E], 30.-31.7.1924, H. Humbert & Perrier de la 
Bathie 2124 (A, P); 24.4.1943, Decary 19116 (P); Valleé de la Betsiboka, en aval de 
Maevatanana [c. 16°50'S, 46°50'E], 1928, H. Humbert & Swingle 4389 (P); Antsalova 
[18°40'S, 44°37'E], Ramaroson 7089 RN (P); Gorges du Manambolo (9e Réserve) [c. 
19°00'S, 45°00'E], 24.-26.10.1932, J. Leandri 436 (G, P, W); Valleé de la Manambolo, Rive 
Droite (Bassin du Mandrare) [c. 19°15'S, 45°00'E], 12.1933, H. Humbert 13006 (B, P), 24.6. 
1930, Decary 7917 (P); Tsimifana, 19°42'S, 44°33'E, 21.3.1971, D. J. Mabberley 754 (K); 
Beroroha [21°40'S, 45°10'E], 1.3.1943, M. R. Decary 18870 (BM 20967, P, S); Ivohibe 
[22°28'S, 46°53'E], 1.1906, d’ Alleizette (L); route Ihosy-Betroka, km 93 de l’Horombe [c. 
23°00'S, 45°30'E], 12.2.1949, R. Capuron 289 SF (P); Beza Mahafaly Reserve near Betioky, 
23°41'S, 44°36'E, P. Phillipson 1839 (P); Beza Mahafaly Reserve, 23°39'S, 44°38'E, 
2.3.1988, L. Sussman 460 (MO); Valleé de l ‘Ianapaly, prés Manera [c. 23°00'S, 44°30'E], 
13.12.1946, H. Humbert 19710 (P); Gorges du Fiherenana, 16.-19.1.1947, H. Humbert 19923 
(P); environs de Tulear: Delta de Fiherenana [23°20'S, 43°40'E], 14.-26.9.1924, Humbert & 
Perrier de la Bathie 2474 (G, P); along Route #10 between Betioky and Ejeda [c. 24°00'S, 
44°30'E], 14.2.1975, T. B. Croat 31260 (P); bassin du Mangoky bei Bevoay [c. 24°12'S, 
45°19'E], 2.11.1959, W. Rauh M1603/59 (M); Berenty Reserve [? 24°34'S, 45°32'E], 
20.4.1991, A. Freidberg & Kaplan 7 (K); région de la Réserve Naturelle Integrale No. 11 
Andohahela, 24°31'S, 46°37'E, 17.9.-30.10.1991, A. Randriamanantena & Durbin 11 (MO); 
Ampany [24°40'S, 44°45'E], 27.11.1932, J. Leandri 645 (BM 21046, P, S); 7.1910, H. 
Perrier de la Bathie 3344 (P); Ifotaka [24°47'S, 46°06'E], 17.11.1938, H. J. Lam & Meeuse 
5452 (BR, L, P); Manasoa Tanosy, 9.1.1913, K. Afzelius (S); 1896, Perrier de la Bathie 83 
(P), Perrier de la Bathie 82 bis (P), M. C. Richard 36, 122 (P). 


5. Pluchea dentex R. Br. Ex Benth.

AUSTRALIA: WESTERN AUSTRALIA: Cape Range Peninsula, entrance to Shothole Canyon, 
22°02'S, 114°04'E, 2.10.1995, Hunger & Kilian 4205 (B, PERTH, etc.); Rocky Pool, 
Gascoyne River, approx. 850 km N of Perth, 24°45'S, 114°08'E, 10.1975, K. F. Kenneally 
4640 (PERTH 759228); Wooramel R. bridge, Perth Carnarvon Rd. [25°45'S, 114°46'E], 
2.9.1959, N. T. Burbidge 6481 (CANB 84374); near Range Bore, 18 km SW of “Lyons 
River” homestead, 24°42'S, 115°10'E, 17.8.1987, K. Newbey 11543 (PERTH 824879); 
Barrow Island, 20°46'S, 115°24'E, 10.1980, R. Buckley 6854, 6911 (PERTH 531731, 
532282), 24.8.1973, W. H. Butler 123 (PERTH 533734); NW Coastal Hwy crossing Cane 
River, c. 22°00'S, 115°36'E, 4.10.1995, Hunger & Kilian 4249 (B, PERTH, etc.); NW Coastal 
Hwy crossing Warramboo Creek, c. 21°40'S, 115°47'E, 4.10.1995, Hunger & Kilian 4252 (B, 
PERTH, etc.); Enderby Island, western end, near Rocky Headland [20°40'S, 116°25'E], 
13.5.1982, B. R. Maslin 5242 (PERTH 452459); 25 miles north of Yalgoo [c. 27°50'S, 
116°40'E], 9.1966, J. R. Knox 660919 (PERTH 532126); 2 km W of Anniversary Bore, 
Jingemarra Station, 27°49'S, 116°47'E, 15.9.1987, R. J. Cranfield 6065 (PERTH 869643); 
Burrup Peninsula, Burrup Range between Withnell and Conzinc Bay, c. 20°33'S, 116°50'E, 
6.10.1995, Hunger & Kilian 4286 (B, PERTH, etc.); Mount Augustus, 24°20'S, 116°51'E, 
24.8.1987, K. Newbey 11683 (PERTH 831107), 9.1971, M. Wittwer S1758 (PERTH 532193); 
15 km E of Namarwell Mt Gould [c. 25°46'S, 117°24'E], 8.1985, A. L. Payne 61 (PERTH 
752754); Roebourne – Wittenoom Rd, near Watthangany Bluff, c. 21°40'S, 117°30'E, 
24.9.1995, Hunger & Kilian 4004 (B, PERTH, etc.); Bally paddock, 2 km N of Rocky Bore, 
Milly Milly Station, 25°47'S, 117°38'E, 24.4.1986, R. J. Cranfield 5329 (PERTH 890375); 
Depuch Island [20°37'S, 117°41'E], 29.5.1962, R. D. Royce 7123 (PERTH 532185); Mt 
Gould Police Station (abandoned), 25°47'S, 117°49'E, 1.5.1986, R. J. Cranfield 5412 (PERTH 
798061); below road crossing at head of Bee Gorge, Hamersley Range NP, 22°30'S, 
118°00'E, 8.8.1973, A. S. Weston 8413 (PERTH 532029); 20.7 miles N of Belele, Eremaean 
Province [27°00'S, 118°00'E], 2.1959, N. H. Speck 939 (CANB 108294); Hammersley Range, 
8.1958, W. D. L. Ride (PERTH 531766); Karijini NP, Hancock Gorge, c. 22°20'S, 118°14'E, 
27.9.1995, Hunger & Kilian 4040 (B, PERTH, etc.); Devels Staircase [22°18'S, 118°17'E], 
25.7.1966, J. V. Blockley 332 (CANB 469610); Karijini NP, Knox Gorge, c. 22°21'S, 
118°20'E, 27.9.1995, Hunger & Kilian 4039 (B, PERTH, etc.); Karijini NP, Kalamina Gorge, 
c. 22°23'S, 118°24'E, 26.9.1995, Hunger & Kilian 4036 (B, PERTH, etc.), Hunger & Kilian 
4038 (B, PERTH, etc.); Wittenoom [22°19'S, 118°21'E], 8./9.1957, S. Elliott 15 (PERTH 
532258), S. Elliott (PERTH 532711); Wittenoom Gorge, 9.1959, K. Stewart (PERTH 
532142); 6 mls S of Wittenoom [22°20'S, 118°21'E], 24.8.1960, A. S. George 1015 (PERTH 
532673); Yampire Gorge, c. 15 km N of Dales Gorge, 22°24'S, 118°24'E, 30.4.1977, R. 
Pullen 10930 (CANB 264026, 264025, MO 4332492); Hillside Woodstock Rd, E of 
Woodstock Homestead, 21°41'S, 119°03'E, Hunger & Kilian 3847 (B, PERTH, etc.), 
21.9.1995, Hunger & Kilian 3854 (B, PERTH, etc.); Roy Hill Rd, shortly E of crossing Port 
Headland – Newman Railway, c. 22°34'S, 119°18'E, 23.9.1995, Hunger & Kilian 3940 (B, 
PERTH, etc.); 36 km N on Bamboo Springs Road [c. 21°45'S, 119°25'E], 15.10.1978, H. 
Demarz 7025 (PERTH 759198); N branch, Gascoyne R., Gt. Northern Hwy. [24°40'S, 
119°30'E], 22.8.1960, A. S. George 956 (PERTH 532231); Booloo Downs Station – Neds 
Gap, [Bulloo Dawns: 24°00'S, 119°35'E], 22.7.1968, J. V. Blockley 771 (PERTH 532037), W. 
E. Blackall (PERTH 532118); Marble Bar Rd between Nullagine and Bonney Downs 
Homestead, 22°00'S, 120°05'E, 22.9.1995, Hunger & Kilian 3879 (B, PERTH, etc.); 9 miles 
N of Nullagine [21°50'S, 120°06'E], 9.1971, R. Mirrington 710918 (PERTH 759236); 2 miles 
N of Nullagine [21°56'S, 120°06'E], 28.10.1970, H. Demarz D2813 (PERTH 532096); 20 
miles N of Mundiwindi [23°30'S, 120°14'E], 20.4.1966, J. V. Blockley 144 (CANB 469611, 
PERTH 532045); Uramurdah Creek, 20 km E of Wiluna, 26°38'S, 120°25'E, 6.9.1982, L. A. 
Craven 7510A (PERTH 856150); Lawlers [28°05'S, 120°32'E], 7.1899, W. V. Fitzgerald 
(PERTH 532150); 10 miles N of Mt Sir Samuel [c. 27°30'S, 120°39'E], 17.10.1945, C. A. 
Gardner 7935 (PERTH 532088); Jig[g]along Depot [23°28'S, 120°44'E], 12.5.1947, R. D. 
Royce 1519 (PERTH 532177); Windich Spring, Canning Stock Route [25°33'S, 120°49'E], 
8.1964, W. H. Butler, R. Aitken & D. Hutchison H.A.61 (PERTH 532002); 18 km N of Mt 
Traine, c. 100 km ENE of Nullagine, 21°46'S, 121°00'E, 6.7.1984, K. Newbey 10358 (PERTH 
532010); Googhenama Creek, on R.P.F. [c. 21°58'S, 121°12'E], 18.5.1947, R. D. Royce 1799 
(PERTH 532061); 15 miles N of Leonora [28°30'S, 121°20'E], 10.9.1953, H. F. & M. 
Bro?ber 1689 (BM 20935); 25 km N of Leonora [28°32'S, 121°20'E], 21.11.1978, H. Demarz 
7248 (PERTH 532169); Yandal, Lake Darlot [27°40'S, 121°35'E], 9.1939, W. E. Blackall 
(PERTH 532053, 532223); Rudall River Region, [22°20'S, 122°05'E], 9.1986, W. G. 
Martinick & Associates (PERTH 1240722); 60 km N of Laverton, [28°00'S, 122°25'E], 
27.8.1968, P. G. Wilson 7319 (PERTH 758744); Minnie Creek, East of Laverton, [28°49'S, 
122°25'E], 1.9.1961, A. S. George 3002 (B); Miss Gibson Hill, SW of Warburton [26°52'S, 
126°20'E], 25.8.1962, A. S. George 4048 (PERTH 759201); Mt Eveline, SE of Warburton 
[26°09'S, 127°07'E], 27.8.1961, A. S. George 2926 (MO 1794611, PERTH 531634); 
Rawlinson Ra., Abercerraga, c. 24°58'S, 128°17'E, 24.7.1974, A. S. George 12138 (PERTH 
532134); base of Schwerin Mural Crescent near Gill Pinacle, 24°53'S, 128°48'E, 17.6.1977, 
A. A. Burbidge & P. J. Fuller 38/77 (PERTH 532207); Ewalinga Rockhole, 20 miles E 
Docker Creek Settlement [Docker Creek: 24°34'S, 128°50'E], 18.9.1969, J. R. Maconochie 
762 (PERTH 531669).
NORTHERN TERRITORY: Hull River, 24°56'S, 129°19'E, 26.8.1973, P. K. Latz 4172 (MO 
2435775); Petermann Ranges Reserve. Chirnside Ck., 25°05'S, 129°38'E, 25.3.1980, T. S. 
Henshall 2889 (CANB 291006); summit of Ayers Rock, [25°21'S, 131°01'E], 30.8.1957, R. 
Schodde 412 (CANB 469620, G, K); Mt Doreen, 22°17'S, 131°17'E, 15.1.1972, C. Dunlop 
2398 (CANB 221379); Mt Sonder, [23°34'S, 132°35'E], 10.9.1958, G. Chippendale (CANB 
161055); Standley Chasm, 35 miles W. Alice Springs, [23°43'S, 133°28'E], 16.10.1957, G. 
Chippendale (CANB 63367), 27.2.1961, H. S. McKee 8658 (K); N. face Mt Gillen 2 miles W 
Alice Springs [23°42'S, 133°40'E], 14.10.1971, P. K. Latz 1807 (PERTH 759147); Stirling 
Creek, near 9 Mile Waterhole, 21°40'S, 133°47'E, 19.3.1985, G. Leach & Smith R 693 
(CANB 359246, MO 4334545); Neutral Junction Station, 21°42'S, 133° 57'E, 4.7.1974, P. K. 
Latz 5610 (K); 9 miles N of Wauchope, Stuart Hwy, 20°33'S, 134°17'E, 20.7.1971, P. K. Latz 
1563 (CANB 220492); Hatches Creek, 20°52'S, 135°12'E, 16.2.1972, C. Dunlop 2501 
(CANB 233306); 11 miles NE of Alexandria Station, [19°06'S, 136°54'E], 22.6.1948, R. A. 
Perry 1475 (CANB 108290, K).
QUEENSLAND: Moongerrie, [Mongogarie?: 28°57'S, 152°56'E], 16.1.1928, P. D. Riddell (K); 
8 km S of Mt Isa [20°52'S, 139°29'E], 3.7.1974, P. Callerenshaw 1171 (K); along railroad 
bank, Jericho [23°34'S, 146°10'E], 1.5.1946, Clemens (A, BM 20959, C, G, K); 15 km S of 
Mt McConnell HS (site 13/6-11), 20°56'S, 146°59'E, 11.6.1992, E. J. Thompson & Sharpe 
CHA59 (K); Mt Slopeaway, near Marlborough, Port Curtis District, [22°51'S, 149°50'E], 
30.8. 1963, N. H. Speck 1749 (CANB 138950).
SOUTH AUSTRALIA: Mt Lindsay, 27°02'S, 129°54'E, 6.8.1962, D. E. Symon 2560 (CANB 
167785, K); c. 18 km NE of Mt Cooperinna, 26°15'S, 130°05'E, 8.9.1978, N. N. Donner 6618 
(K); Gammon Range NP, Flood plain of Italowie Creek, 30°30'S, 139°09'E, 3.10.1981, W. 
Bushman 19 (AD 98206108), Italowie Gorge, J. B. Cleland (AD 97413394); Paralana 
Springs, [30°11'S, 139°27'E], 24.8.1968, D. E. Symon 6008 (CANB 196990); Creekline +/- 6 
miles W of S. A.-Q. border fence, N of the Cooper, between Innamincka and Nappamerrie 
Stn., [27°40'S 140°54'E], 19.8.1968, D. E. Symon 5795 (B, CANB 358666, K).


6. Pluchea dioscoridis (L.) DC.

CHAD: Kanem, Kinémokadi, 4.7.1935, Murat 153 (P 11657); Lac Tchad et Kanem 
11.10.1903, Chevalier 10227 (B, P 11644); Tchadsee, Gebiet Schari, 9.7.1899, Chevalier 
1159 (B); am Selep [13°22'N, 16°46'E], 19.7.1964, Gaston 5218 (P 11647).

LIBYA: Muh, bei Ain Hesne ?, 1.2.1874, G. Rohlfs Expedition 772 (LD).

EGYPT: Nubia, El-Dakhla, Ballana [22°16'N, 31°34'E], 9.1.1964, L. Boulos (LD); Nile Valley, 
Kom Ombo, N of Aswan, [24°26'N, 32°57'E], 19.4.1967, Gun Romée (LD); El Faiyum 
[29°19'N, 30°50'E], 15.2.1974, Amal Amin, Mahid, Sisi & Abbel Aziz (LD); Suez [29°58'N, 
32°33'E], 9.5.1904, A. Kneucker 103 (B); Nile Valley, Cairo, [30°03'N, 31°15'E], 4.5.1967, 
Gun Romée (LD), 1822, Sieber (GOET), 1846, Sieber (GOET), 6.1876, Sieber (C), Sieber 
(BR), 31.3.1881, Letourneux (A), 8.4.1848, Herb. L. Kralik (A), Boulaq el Dakrour, Giza, 
12.6.1971, Soliman Sisi (LD), Saft El Iaban, 20.8.1974, Soliman Sisi (LD), in palmetis ad El-
Marg., 27/28.4.1908, J. Bornmüller 10736 (B), 4.5.1908, J. Bornmüller 10735 (B), 25.8.1864, 
C. Schweinfurth 256 (BM 21114, 21115, W); El Birka [30°09'N, 31°22'E], 10.10.1944, P. D. 
Davis 7258 (E); Inter Ismailija et Sues, prope Fayid adaquar, [c. 30°18'N, 32°16'E], 
26.5.1908, J. Bornmüller 10734 (B, E, HBG); Abu [al] Matamir western Delta [30°54'N, 
30°10'E], 28.5.1967, V. Täckholm & I. Elsayed & M. el Mahdi (LD); Nile Valley Tall 
Hammad, 60 km S from Cairo, 22.4.1967, Gun Romée (LD); Gabbari [31°11'N, 29°53'E], 
3.6.1877, A. ? (LD); Alexandria [Al Iskandariyah: 31°12'N, 29°54'E, J. Pfund (C), 1.1919, G. 
F. Scott Elliot 3223 (BM 21118); Menzaleh Delta [31°13'N, 32°07'E], 13.7.1887, 
Schweinfurth (BM 21114); Rosetta [31°24'N, 30°25'E], E. D. Clarke (BM 20956), 24.9.1944, 
P. H. Davis 7184 (E); prope Atfe Egypti inf., 30.11.1834, Schimper & Wiest (M, W).

SUDAN: Sennar [13°31'N, 33°38'E], 1839, ? 462 (GOET); below Guldu, on west side of Marra 
Mts. [c. 13°45'N, 24°30'E], 2.1.1934, J. E. Dandy 31 (BM 20964).

ETHIOPIA: Tigre [13°14'N, 39°21'E], 1.11.1862, Schimper 697 (BM 21108, C); about 35 km S 
of Addis Ababa, [8°50'N, 38°42'E], 1.4.1965, G. J. H. Amshoff 6091 (B), 1.4.1965, de Wilde 
6091 (B, BR, C); Wello Region, Ambasel Awraja Anabe Forest [c. 11°35'N, 40°00'E], 
28.10.1987, T. Mesfin 6515 (C); Errer River, about 20 km on the road from Harar to Jijiga, 
[9°20'N, 42°20'E], 31.12.1968, J. J. F. E. Wilde 4311 (B, BR, C); Debre Zeit, ILCA 
Experimental Field Station [c. 9°00'N, 39°00'E], 12.10.1976, L. Boulos 9888 (BR); Gemu 
Gofa: 80 km W of Hamer Koke [5°45'N, 37°00'E], 10.8.1969, M. G. & S. B. Gilbert 1539 
(C); about 10 km S of Nefasit [15°15'N, 39°02'E], 2.11.1969, J. J. F. E. de Wilde 4518 (B, 
BR); Senaar, Kotschy 155 (BM, GOET, M, P 11526, W); in M’has, 8.1872, Hildebrandt 410 
(W).

ERITREA: Baria [Baia?: 13°39'N, 42°07'E], 9.1.1906, A. Pappi 6812 (C); Hamasen, lungo il 
torrente Fil-Fil [15°37'N, 38°58'E], 23.5.1902, A. Pappi 5444 (BM 21077, HBG, W); Eritrea-
Beni Amer: Tr· Scetel e Carajai? [15°50'N, 37°30'E], 8.3.1906, A. Pappi 6956 (A, BM 21081, 
G, P 11528); alle Porte del Diavolo, 1.1.1902, A. Pappi 5002 (BM 21079); Eritrea-Bari·: 
lungo il torrente Sandacca, 28.1.1906, A. Pappi 6821 (BM 21080, G); Valle Ant, 1.3.1892, 
Schweinfurth & Riva 678 (G).

DJIBOUTI: Source Ouéah, 25.3.1938, M. Aubert de la Rue (P 11521).

CONGO: Lac Edouard, baie de Vitshumbi [c. 0°25'S, 29°30'E], 3.1.1953, Van der Ben 11 
(BR); Beni [0°29'N, 29°29'E], 1.8.1914, Bequaert 5146 (BR); Kisangani [0°33'N, 25°14'E], 
20.11.1982, S. Lisowski 86038 (BR); Ruindi [c. 0°47'S, 29°17'E], 1937, J. Lebrun 8372 (BR); 
May yo Moto[0°54'S, 29°21'E], 1937, J. Lebrun 7640 (BR); Rutshuru [1°11'S, 29°26'E], 
16.11.1971, C. Evrard 6825 (BR); Rive N. du lac Kivu á Sake [c. 1°34'S, 29°02'E], 1929, 
Humbert 8425 bis (BR, P 11661), Lushadu, bords lac Kivu, 20.3.1960, Hendrickx 7737 (BR); 
lac Kivu, Ibitagata [c. 2°00'S, 29°10'E], 20.8.1953, Van der Ben 719 (BR); Vallée Rwindi, 
4.4.1959, Keremera 35 (BR), 3.7.1956, G. F. de Witte 13240 (BR); Kabare, 18.8.1919, 
Bequaer 5314 (BR); PNA. Sect. Kasindi (lac), 14.11.1949, de Wilde 401 (BR); Kokondo, 
28°52'E, 2°15'S, 21.11.1948, Hendrickx 5647 (BR).

BURUNDI: KABURANTWA-TERRITOIRE: Bubanza, [3°05'S, 29°22'E], 30.10.1970, J. Lewalle 
4883 (BR, C, G); Plaine de la Rusizi au km 30 de la Route Bujumbura-Cibitoke, [c. 3°22'S, 
29°19'E], 30.5.1975, J. Lambinon 75/57 (LD); Delta de la Ruzizi, 22.4.1980, J. Lambinon 
80/433 (BR); Bujumbura, 3°22’S, 29°16'E, 5.3.1977, M. Reekmans 5784 (BR); Bujumbura, 
3°32'S, 29°21'E, 12.5.1982, M. Reekmans 11177 (BR).

KENYA: Tsavo East Nat. Park [c. 3°04'S, 39°10'E], 26.2.1977, S. Hooper & C. Townsend 
1086 (C).

ANGOLA: Bengo inundatis a Prata [9°59'S, 14°35'E], 9.1857, Welwitsch 3926 (BM 21165, G, 
P 11553); Distr. Pungo Andongo, 1857, Welwitsch 3925 (BM 21164, P 11552).

SAUDI ARABIA: road to Najrahn, 50 km N of Dharan [17°40'N 43°30'E], 2.3.1981, D. Hillerat 
75 (BM 21009); Najran [17°30'N, 44°10'E], 1.1979, S. Chaudhary 266 (E); W Al Ashiba 
[17°43'N, 44°25'E], 15.4.1950, K. M. Guichard KG/11/Tihama (BM 20995); Mahail, 
18°30'N, 42°00'E, 24.12.1971, G. Popov 71/323 (BM 21090); Wadi Adamah, 19°54'N, 
41°55'E, 27.10.1969, J. P. Mandaville 2652 (BM 21062); Al-Baha, Wadi Habaqa, S of Al-
Baha [20°00'N, 41°30'E], 10.10.1985, A. A. Fayed 1183 (B, M); Wadi Aridah, 20°37'N, 
47°17'E, 31.10.1969, J. P. Mandaville 2692 (BM 21063); Thaqif highlands, 20°45'N, 
40°50'E, 8.12.1946, D. Vesey-Fitzgerald (BM 21154); Al-Taif, Al-Hada region, about 18 km 
from Al-Taif [21°16'N, 40°24'E], 22.3.1985, A. A. Fayed 1121 (B); southern Tihama foothills 
[c. 22°00'N, 40°00'E], 2.1971, G. Popov 71/49 (BM 21089); Darin village, Tarut Island, 
[26°35'N, 50°05'E], 2.10.1987, J. P. Mandaville, Jr. 8658 (E); Arabia, 1820-1826, Ehrenberg 
(BR); Wadi Said near Said, 13.3.1952, G. Popov, L. Tillin & H. B. Gilliland (BM 21092); 
Wadi Hajar, W of Maifa, 24.8.1949, K. M. Guichard KG/HAD/120 (BM 20996); W Hagir, 
16.3.1952, J. D. Tothill 63 (BM 21149).

UNITED ARAB EMIRATES: Abu Dhabi [24°28'N, 54°25'E], 29.5.1982, R. Western 144 (E).

OMAN: Dhofar, Jabel Qamar, 16°52'N, 53°45'E, 9.10.1998, P. Hein 5561 (herb. P. Hein), 
9.10.1998, N. Kilian & P. Hein NK 5603 (B); Muscat – Qurm, 23°37'N, 58°28'E, 1.10.1998, 
P. Hein 5129 (Herb. P. Hein), 1.10.1998, N. Kilian & P. Hein NK 5260 (B).

YEMEN: Wadi Arafat, on Taiz to Turbah road [c. 13°00'N, 44°00'E], 16.3.1984, A. G. Miller 
& R. A. King 5194 (E); Al Lasab, SE of Ta’izz on Salah-Adanan road, 28.7.1983, K. Gordon 
127 (E); Oberlauf des Wadi Labtah – südl. Beckenrand von At Tur [c. 13°00'N, 44°20'E], 
11.3.1982, Müller-Hohenstein & Deil 262 (E); S facing escarpment into Wadi Urbil at the S 
edge of At Turbah, 13°12'N, 44°07'E, 1.4.1997, N. Kilian & P. Hein  NK 4717 (B); Wadi 
Sara, 4 km W of Qamis al Beni Saad [c. 13°27'N, 45°30'E], 30.3.1981, A. G. Miller & D. G. 
Long 3326 (E); Jabal Habaschi, slopes with terraces NE of the village Yafrus, 13°28'N, 
43°56'E, 29.3.1997, N. Kilian, P. Hein & M. Smalla NK 4657 (B); Lahij Governorate, Wadi 
Fare’aa, 8 km E Habil Jabir [c. 13°31'N, 44°56'E], 24.12.1987, A. K. Rowaished, W. A. Saeed 
& M. A. Hussein (BM 21101); Ta’izz [13°34'N, 44°02'E], dry Wadi bed north of town, 
22.12.1937, H. Scott & E. B. Britton 349 (BM 21117); Wadi Barakan between Taiz and 
Turba, 11.8.1975, J. R. I. Wood Y/15/514 (BM 21175); on route from Hays to Udayn, at 
Sheehsly Muzahin [c. 13°55'N, 43°30'E], 22.9.1983, K. Gordon 269 (E); Hammam Ali 
[14°39'N, 44°08'E], 28.8.1937, C. Rathjens 37/25 (BM 21097); km 90 Hodeidah-San’a road 
[c. 15°00'N, 44°00'E], 20.9.1962, G. Popov 8/39 (BM 21085); am Bach im Wadi Dhar bei 
Sanaa [c. 15°21'N, 44°12'E], 14.10.1982, Müller-Hohenstein & Deil 832 (E); 47 km N Huth 
nahe der Stra?e nach Sadah [c. 17°00'N, 43°58'E], 2.10.1981, D. Podlech 36194 (G, M).

JORDAN / PALESTINE: Jericho [31°51'N, 35°27'E], Ball 967 (C); Eliasquelle bei Jericho, 
26.5.1904, A. Kneucker 418 (B), 30.3.1897, J. Bornmüller 827 (B), 1863-64, B. T. Lowne 
(BM 21052, 21053), 31.5.1882, Herb. Postian. (BM 21094); Transjordania, Wadi Nimsin 
[31°54'N, 35°38'E], 5.5.1932, G. Samuelsson 821 (B, LD); Wadi el Kelt, Wüste Juda, 3.1930, 
H. Amsel (B); Jordanthal, Palaestina, 4.1900, Warburg (B); 26.4.1886, Herb. Postian. (BM 
21093); Anjeh river, 7.12.1912, F. S. Meyers & J. E. Dinsmore 34174 (E); Ain et Tabigha 
[?Tabib, Ayn: 31°12'N, 35°39'E], 19.7.1902, F. Meyers 174 (E); Dead Sea, Hunting Aero 
Survey, 10.5.1955, ? 201 (E); Wadi-ul-Kilt, 24.9.1913, J. E. Dinsmore B6174 (G).

ISRAEL: Ein Gedi [31°27'N, 35°23'E], 30.6.1984, K. Lind (C); Nahal Ha’ Ammud, östlich von 
Huqoq, nördlich von Tiberias [32°48'N, 35°32'E], 13.5.1988, S. Rilke 1345 (B).

SYRIA: Hemmé [32°41'N, 35°40'E], 21.1.1952, H. Pabot (G); Jardin a gauche de la route de 
Megg a pres, Damas [Damascus: 33°30'N, 36°19'E], ? 1278 (A, B, BR, C); ?al el Kadi, infra 
Banias [33°14'N, 35°41'E], 2.1.1884, E. Peyron (G).

LEBANON: Beyrouth [33°52'S, 35°30'E], Blanche (W).

IRAN: Bender Abbas (beim Dorf Sarchem), 4.4.1977, Manutschehri 996 (GOET).

IRAQ: Masharra, Amara [31°50'N, 47°10'E], 3.5.1918, W. E. Evans (E).


7. Pluchea dunlopii Hunger

AUSTRALIA: Western Australia: c. 30 km SW of Learmonth, Yardie Creek, 22°24'S, 113°45'E, 
1.10.1995, Hunger & Kilian 4157 (B, PERTH, etc.); near Tracking Station, Carnarvon 
[24°51'S, 113°45'E], 28.10.1963, Fairace & Lullfitz L2813 (PERTH 532967); 12 km S 
Carnarvon [24°57'S, 113°45'E], 5.10.1984, H. Demarz 10304 (PERTH 533580); 10 km S 
Carnarvon [24°56'S, 113°45'E], 8.12.1982, H. Demarz 9516 (PERTH 887765); Yardie Creek, 
22°19'S, 113°49'E, 6.9.1970, A. S. George 10294 (PERTH 533785); 240 miles N of Geraldton 
[25°30'S, 114°00'E], 24.5.1962, T. E. H. Alpin 1514 (PERTH 528404); 72 km E on the 
Gascoyne Junction road from its junction with the NW Coastal Highway to Carnarvon [c. 
24°50'S, 114°00'E], 21.9.1982, N. F. Norris 914 (PERTH 533009); 70 km S Bullara turn off 
[c. 23°30'S, 114°02'E], 10.8.1976, Wittwer 1794 (PERTH 533629); 76 km E of Carnarvon, 
24°49'S, 114°25'E, 27.9.1987, P. G. Wilson 12673 (CANB 469607, PERTH 864919); 
Minderoo, Ashburton River [21°59'S, 115°04'E], 11.10.1905, A. Morrison (PERTH 533637, 
532908); Barrow Island, 20°46'S, 115°24'E, 10.1980, R. Buckley 6958 (CANB 341392); near 
M46 lease, 9.10.1990, M. White MRW057 (PERTH 1390066); Roebourne Wittenoom road, c. 
10 km N of Wittenoom, 22°10'S, 118°20'E, 24.9.1995, Hunger & Kilian 3977 (B, PERTH, 
etc.); c. 12 km by road W of Roy Hill on Wittenoom road, 22°41'S, 119°53'E, 17.8.1977, E. 
N. S. Jackson 2907 (AD 97744610); 133 km E of Wiluna on the road to Warburton, 26°34'S, 
121°25'E, 5.9.1984, P. G. Wilson 11936 (PERTH 533602); Billiluna Station [Homestead?] E 
Kimberley [19°33'S, 127°40'E,], 1992, T. Handasyde 33 (PERTH 3171469).
NORTHERN TERRITORY: 25.5 miles NW Mt Doreen HS, 21°50'S, 131°10'E, 1.5.1958, G. 
Chippendale (CANB 90975); Cockatoo Creek c. 255 km NW of Alice Springs [22°20'S, 
131°30'E,], 13.8.1931, J. B. Cleland (AD 95835052); Haast’s Bluff Reserve [c. 23°20'S, 
131°56'E], 5.9.1957, J. B. Cleland (AD 966060477); Mount Chapple Bore c. 135 km NW of 
Alice Springs [23°17'S, 132°57'E], 31.8.1951, J. B. Cleland (AD 95835053); Numagalong 
H/S [20°49'S, 133°33'E], 26.8.1965, D. Nelson & R. Swinbourne (PERTH 533866, AD 
96550466); Macdonnell Ranges, 23°38'S, 133°50'E, 19.10.1979, D. J. Nelson 2486 (BM 
21074, G); C.S.I.R.O. Field Station, Alice Springs, 23°46'S, 133°52'E, 5.8.1980, D. J. Nelson 
2499 (CANB 469612); Ooraminna carvings area C.S., 24°00'S, 133°57'E, 16.10.1975, P. K. 
Latz 6318 (AD 98343178); Stuart Highway between Wauchope and Tennant Creek, 20°13'S, 
134°13'E, 13.8.1978, W. R. Barker 2789 (AD 97921059); Bonny Creek 80,47 km S of 
Tennant Creek (Bonny Creek is 19°33'S, 134°14'E), 23.8.1966, N. Walker (AD 97817428); 
Tomahawk Swamp, Mt Skinner Station [22°08'S, 134°32'E], 14.7.1971, D. J. Nelson 2127 
(AD 97152259, CANB 220565); Ringwood Station, 23°54'S, 134°59'E, 4.10.1973, P. K. Latz 
4439 (CANB 469615); c. 255 km NE of Alice Springs [22°25'S, 135°08'E], 30.8.1933, E. H. 
Isling (AD 97413207, 97413246); NW Simpson Desert, 23°33'S, 136°16'E, 2.11.1973, P. K. 
Latz 4424 (AD 97615069, CANB 469605); on Barkly Hwy 70 km NW of Barry Caves, 
20°20'S, 137°35'E, 1.9.1981, T. & J. Whaite 3927 (MO 4332475).
NEW SOUTH WALES: W shore of Fort Grey basin (Pinaroo “Lake”) 29°05'S, 141°12'E, 
8.9.1981, W. Greuter 18559 (B); Frome swamp, 75 km NW, of Tibooburra, Sturt NP, 
15.6.1979, 29°10'S, 141°18'E, K. Paijmans 3142 (CANB 286376), 11.6.1979, K. Paijmans 
3454 (CANB 292333); “Waka”, SE of Fort Grey, 29°16'S, 141°21'E, 1.11.1972, P. L. 
Milthorpe 887 (AD 97450109); c. 65 km W of Wanaaring along the Tibooburra road, 
29°43'S, 143°26'E, 30.10.1986, P. S. Short 3008 (AD 98707423).
SOUTH AUSTRALIA: Near Kalladeina Bore, 27°42'S, 139°07'E, 2.10.1989, H. M. Lee 540 (AD 
99110390); 64 km S of Birdsville, 26°25'S, 139°33'E, 25.8.1975, J. Z. Weber 4646 (AD 
97537272); 15 km NW Moomba, 28°01'S, 140°05'E, 4.9.1987, J. Reid 1202 (AD 98742077); 
Coopers Creek, c. 85 km NW of Innamincka, 27°8' 10'S, 148°08'E [writing error, it must be 
140°08'], 20.8.1975, J. Z. Weber 4602 (AD 97539217, W); S of Coongie Lake, 27°11'S, 
140°09'E, 21.8.1975, N. N. Donner 5242 (AD 97541004); random site 41.2 km E Cuttaberrie 
Lake, 27°17'S, 140°15'E, 10.9.1987, J. Reid 1155 (AD 98742068); Merty Merty flood plain, S 
of Innamincka, halfway on Strzelecki Creek [c. 27°40'S 140°20'E], 25.5.1957, from S. A. 
Pastoral Board 157 (AD 97636328); Big Lake Moonba, 28°20'S, 140°21'E, 30.9.1983, L. 
Alexander 2223 (AD 98516253); Mudcarnie Well, 26°52'S, 140°35'E, 7.11.1983, L. 
Alexander 2385 (AD 98439320); Coongie sandhills, 27°05'S, 140°36'E, 16.10.1986, P. E. 
Conrick 1957 (AD 98647410); Innamincka, 27°44'S, 140°46'E, 19.9.1959, from S. A. 
Pastoral Board (AD 97629334, 97906020), 17.10.1963 (AD 97629598), 18.10.1963, (AD 
97916049, 98025121), 19.9.1959, J. Reid (AD 96051004, 98671529), 15.10.1963, W. S. Reid 
(AD 98671531); No. 1 Bore track from Innamincka, 19.9.1959, W. S. Reid (AD 96139080); 
30 km E of Toolache Waterhole, 28°15'S, 140°53'E, 11.5.1988, Reid & Gillen 6 (AD 
98829330); 7 km NNE of Candradecka Dam, 27°13'S, 140°52'E, 1.6.1976, L. D. Williams 
8284 (AD 98424800); c. 30 km E of Cordillo Downs, c. 4 km W of Arrabury Homestead, 
26°46'S, 140°59'E, 28.8.1975, N. N. Donner 5288 (AD 97540306).
QUEENSLAND: 70 km NE of Nappa Merrie HS, 27°15'S, 141°36'E, 12.9.1984, F. J. Badman 
1531 (AD 98447096); 14,5 km S of Listowel Valley turn off on Blackall to Adavale road [c. 
25°30'S, 145°06'E], 23.9.1984, M. E. Ballingall 1601 (AD 98522102, CANB 373143); 
Belyando R., near “Moray Downs” Station, [21°57'S, 146°38'E], 3.8.1964, L. G. Adams 1226 
(CANB 152195); Rockhampton [23°22'S, 150°32'E], Thozet (K), F. Mueller (C).
NOT LOCATED: Nerkool Creek, F. Mueller (A); Broad Sound, 18.9.1802, R. Brown 2100 (BM 
20938, K).


8. Pluchea eupatorioides Kurz

THAILAND: Kaun San, Chaiyabhum, [15°46'N, 101°55'E], 18.2.1931, A. F. G. Kerr 20180 
(BM 21036); MeKor, Raheng [Tak: 16°52'N, 99°08'E], 17.3.1915, K. Winih 222 (K, BM 
21171); Maetak [c. 17°00'N, 100°00'E], 3.3.1958, Th. Sørensen, K. Larsen & B. Hansen 1875 
(C, K, L); Lampang, Ban Me Ta [18°08'N, 99°31'E], 18.2.1922, K. Winit 690 (K); Meh Kung 
(near Doi Saket), [18°51'N 99°11'E], 1.3.1910, A. F. G. Kerr 1021 (BM 21028, K); Muang 
Pai [19°19'N, 98°27'E], 1921, Kerr 5508 (B, BM 21031, 21032); Thong Ma Haw to Keng 
Khaep, 27.1.1962, K. Larsen 9320 (C); Ban Salü, 11.2.1905, Hosseus 380 (M, P).

CHINA: Yunnan [25°00'N, 102°00'E], A. Henry (K).

VIETNAM: Prov. Sonla, canton de Muong Mua [21°12'N, 103°58'E], 5.1927, E. Colani 4046 
(K); ?on Cuang a Cua Rao [c. 19°17'N, 104°26'E], Province de Vinh, 4.1931, A. Pételot 2048 
(A).

LAOS: Bassin du Sé-Moun (Laos méridional), 1875-77, Harmand (K).


9. Pluchea ferdinandi-muelleri Domin

AUSTRALIA: WESTERN AUSTRALIA: Between Naroo and Nanutara, Ashburton River 
[Nanutarra: 22°32'S, 115°30'E], 3.10.1905, A. Morrison (A, PERTH 532827); 9 miles S of 
Port Hedland, 20°25'S, 118°35'E, 10. 1946, C. M. Donald (CANB 13663); Carnarvon 
[24°51'S, 113°45'E], G. B. Barnett (PERTH 532797); E of Learmonth airfield, 22°17'S, 
114°07'E, 8.9.1970, A. S. George 10345 (PERTH 532991); Barrow Island, 20°46'S, 115°24'E, 
10.1980, R. Buckley 6852 (PERTH 532940); Cleaverville track, between Karratha and 
Roebourne [c. 20°45'S, 117°00'E], 7.10.1982, coll. ignot. 335 (CANB 353129, 353130, 
PERTH 532983); North West Coastal Hwy, c. 7 km E of Whim Creek, 20°51'S, 117°59'E, 
19.9.1995, Hunger & Kilian 3762 (B, PERTH, etc.); Peeawah R., NW Coastal Highway [c. 
20°50'S, 118°10'E], 27.8.1960, A. S. George 1113 (PERTH 532959); Boodarie [Homestead?] 
Station, South Hedland [20°24'S, 118°28'E], 22.11.1984, J. Stretch (PERTH 533599); 8 miles 
S from Port Hedland [20°34'S, 118°36'E], 7.9.1946, C. A. Gardner 8060 (PERTH 532851); 
Great Northern Hwy, c. 80 km S of Port Hedland, 21°07'S, 118°42'E, 20.9.1995, Hunger & 
Kilian 3808 (B, PERTH, etc.); Great Northern Hwy, c. 150 km S of Port Hedland, 21°38'S, 
118°48'E, 21.9.1995, Hunger & Kilian 3828 (B, PERTH, etc.); between Hillside and 
Woodstock Homestead 21°41'S, 119°03'E, 21.9.1995, Hunger & Kilian 3852 (B, PERTH, 
etc.); near Woodstock Homestead, 21°44'S, 119°17'E, 21.9.1995, Hunger & Kilian 3839 (B, 
PERTH, etc.); between Hillside and Woodstock Homestead, 21°48'S, 119°29'E, 21.9.1995, 
Hunger & Kilian 3860 (B, PERTH, etc.); Bamboo Springs road [22°00'S, 119°25'E], 
27.10.1970, H. Demarz 2811 (PERTH 533661); 14 miles N of Marble Bar [21°16'S, 
119°45'E], 9.1971, R. Mirrington 710912 (PERTH 532843); c. 10 km S of Nullagine, 
22°00'S, 120°05'E, 21.9.1995, Hunger & Kilian 3877 (B, PERTH, etc.); Wallal [Wallal 
Downs: 19°50'S, 120°45'E], 9.1922, Herb. M. A. Evershed (BM 20976); 124 miles SW of 
Anna Plains, SW of Broome [19°50'S, 120°49'E], 9.8.1965, A. C. Beauglehole 11329 (CANB 
343310, PERTH 533068); Mandora Soak, NE of Radi Hills, 19°45'S, 121°26'E, 13.8.1983, N. 
McKenzie MS-06 (PERTH 859397); Dampier Peninsula, 17°22'S, 122°09'E, 8.10.1984, P. R. 
Foulkes 16 (PERTH 533149); Point Coulomb Nature Reserve, 17°16'S, 122°11'E, 8.8.1992, 
A. A. Mitchell 2652 (CANB 464431, PERTH 2632667); Roebuck Bay [c. 18°00'S, 122°15'E], 
1890, J. Tepper (PERTH 1625845); Little Sandy Desert, 23°06'S, 123°20'E, 2.5.1979, A. S. 
Mitchell 935 (PERTH 532762); Great Sandy Desert, 19°40'S, 123°21'E, 10.8.1977, A. S. 
George 14745 (PERTH 532916); Gogo Station, [18°17'S, 125°35'E], 5.10.1952, K. Fitzgerald 
(PERTH 532835); Nimbi Caves area, Lawford Range, 18°42'S, 126°00'E, 3.11.1986, C. Done 
746 (PERTH 2246546); 76 miles W of Margaret River, SSE of Fitzroy Crossing [c. 17°30'S, 
126°45'E], 1.8.1965, A. C. Beanglehole 11150 (PERTH 533076); Gordon Downs Station 
[18°43'S, 128°33'E,], 13.9.1950, R. D. Royce 3330 (PERTH 533084).
NORTHERN TERRITORY: 71 miles WNW Tanami [19°35'S, 129°10'E], 13.9.1971, C. R. Dunlop 
2338 (AD 97422240, 98671532, CANB 237562); 1 km SE Supplejack Station H/S, 19°21'S, 
129°58'E, 4.10.1978, D. J. Nelson 2455 (CANB 286733, 286435); c. 25 km by road NW of 
The Granites, 20°20'S, 130°10'E, 20.8.1978, N. N. Donner 6311 (AD 97849486); The 
Granites [20°38'S, 130°22'E], 8.1936, J. B. Cleland (AD 95835057); about 30 m[iles] W Mt 
Doreen [22°10'S, 131°05'E], 24.6.1957, G. Chippendale (AD 96549309, PERTH 533726); 
Mount Palmer [c. 23°30'S, 131°10'E], 26.8.1956, J. B. Cleland (AD 95835058); Mount 
Liebig [23°16'S, 131°18'E], 14.8.1932, J. B. Cleland (AD 95835055); Brooke’s Soak [c. 
22°00'S, 131°30'E], 15/16.8. 1931, J. B. Cleland (AD 95835056); c. 13 km W of Yuendumu 
[Mission] [22°20'S, 131°37'E], 18.8.1951, J. B. Cleland (AD 95835054); Mt Wedge Station 
[23°50'S, 131°50'E], 25.8.1957, J. B. Cleland (AD 966060476); 8 miles E Haast Bluff 
Settlement [23°20'S, 132°03'E], 24.8.1956, G. Chippendale (AD 95910077); 1 miles S of Mt 
Wedge Station [22°45'E, 132°09'E], 29.9.1956, M. Lazarides 6063 (AD 96003025, CANB 
108295, PERTH 533718); 42 miles NW Willowra H. S. [c. 20°55'S, 132°25'E], 30.7.1958, G. 
Chippendale (AD 966030152, CANB 214955); 35 miles S of Napperby Station [22°50'S, 
132°35'E], 28.9.1956, M. Lazarides 6084 (AD 96003023, B, CANB 108296, PERTH 
533696); Narwietooma Station, 23°08'S, 132°36'E, 30.1.1976, T. S. Henshall 1205 (CANB 
469606); Mt Giles, 23°39'S, 132°50'E, 21.9.1976, P. K. Latz 6629 (CANB 267410, MO 
2433988); Prowse’s Gap [22°34'S, 133°20'E,], 1954, T. R. N. Lothian 382/54 (AD 96218047); 
Alice Springs, 23°42'S, 133°52'E, 1954, T. R. N. Lothian (AD 97648051); NW Simpson 
Desert, 23°52'S, 136°28'E, 1.10.1973, P. K. Latz 4403 (AD 97511104, CANB 469613); 6.1 
miles N of Georgina Downs H. S [c. 21°00'S, 137°39'E], 1.10.1957, G. Chippendale 3804 
(AD 95951155, CANB 70038, PERTH 533688).


10. Pluchea glutinosa Balf.

YEMEN: Socotra, [near Gallonsir stout], Balfour 223 (K[b], OXF).


11. Pluchea grevei (Drake) Humbert

MADAGASCAR: Morondava [20°19'S, 44°17'E], 7.1891, Douliot (P), 11.9.1924, H. Humbert & 
Perrier de la Bathie 2380 (A, P); environs de Morondava [20°22'S, 44°22'E], 9.1956, J. 
Bosser 9917 (P); Fort de Marofandelia entre Morondava et la Tsiribihina [?], 10.1933, H. 
Humbert 11427 (P, S); Itampolové, 60 miles N of Tulear [22°15'S, 43°16'E], 18.6.1929, 
White (BM 21168); 8 km N of Tulear on road to Morombe [c. 23°10'S, 43°40'E], 5.2.1975, T. 
B. Croat 30776 (P); environs de Tulear: Delta de Fiherenana [23°20'S, 43°40'E], 14.-
26.9.1929, H. Humbert & Perrier de la Bathie 2401 (G, P), 6.8.1928, H. Humbert & Swingle 
5180 (B, BM, P), 15.10.1970, M. Keraudren-Aymonin & al. 24698 (P), basse vallée du 
Fiherenana, 11.1933, H. Humbert 11582 (P), 15.10.1970, M. Keraudren-Aymonin & al. 
24694 (P); Tulear [23°20'S, 43°41'E], 11.-1.1868/69, Grandidier (P), 8.11.1967, L. Bernardi 
11316 (G, L), 1910, H. Perrier de la Bathie 3346 (P), 1840, M. Richard (P), 5.11.1959, W. 
Rauh 1101/59 (M), Region de “la Table”, 1.1922, G. Petit 15 (P); côte Mahafaly, Besambaly 
prés Beheloka [23°30'S, 43°30'E], 6.1910, H. Perrier de la Bathie 3342 (P); Tongobory 
(Onilahy) [23°30'S, 44°20'E], 8.1919, H. Perrier de la Bathie 12767 (P); Estuaire de 
l’Onilahy a St. Augustin [23°31'S, 43°46'E], 21.-23.1.1947, H. Humbert 19969 (P); Distr. 
Betioky, Soalary [23°35'S, 43°43'E], 25.11.1952, Ravelonanahary 4294-RN (P); Southern 
Domain, 40 km NE of Betioky, 23°39'S, 44°38'E, 23.7.1987, L. Sussman 221 (MO), 
12.8.1987, L. Sussman 246 (MO), 1.9.1987, L. Sussman 259 (MO); Anakao [23°40'S, 
43°40'E], 7.10.1921, H. Poisson 336 (P); basin inférieur de l’Onilahy, environs de Betioky [c. 
23°40'S, 44°20'E], 11.-15.8.1928, H. Humbert & Swingle 5248 (P); environs du Lac 
Tsimanampetsotsa [24°10'S, 43°54'E], 13.-16.2.1947, H. Humbert 20246 (P); Mahene 
[24°50', 44°55'E], 12.12.1917, R. Decary (G, P); between Ampanihy and Beloha [c. 24°50'S, 
45°00'E], 16.2.1975, T. B. Croat 31461 (MO, P); Antanimora, prov. Fort-Dauphin [24°50'S, 
45°45'E], 9.8.1924, M. Decary 2975 (BM 20965, C, P, S), 16.7.1926, M. Decary 4350 (G, P, 
S), 20.7.1926, M. R. Decary 4529 (P);  from Antanimora to Beraketa, 21.2.1975, T. B. Croat 
32040 (MO, P), 32045 (P), 27.1.1990, N. Dumetz 1262 (MO), 18.10.1970, F. R. Fosberg 
52527 (MO); environs du Lac Manampetsotsa, delta de Linta [24°55'S, 44°00'E], 1928, H. 
Humbert & Swingle 5304 (A, P), 5335 (A, P), 5425 bis (A, P), 5477 bis (G, P); Androka 
[24°55'S, 44°10'E], 6.1910, H. Perrier de la Bathie 3343 (P); about 60 road-km W of Fort-
Dauphin (Tôlanaro), 24°59'S, 46°33'E, 24.5.1991, J. L. Zarucchi & al. 7483 (K, P); 
Ambovombe, vallée du Mandrare a Amboasary [25°01'S, 46°23'E], 1.10.1931, R. Decary 
9250 (BM 20966, P); Ambovombe [25°10'S, 46°06'E], 17.4.1924, M. R. Decary 2592 (P), 
3983 (L), 26.2.1931, M. R. Decary 8562 (P, W); Lahimanara, 8.6.1931, R. Decary 8990 (P); 
lac Anongy, Dist. Ambovombe [25°10'S, 46°32'E], 11.6.1924, M. R. Decary 2856 (P); Fort-
Dauphin, [Cap?] Andrahomana [25°15'S, 46°40'E], 22.9.1932, R. Decary 10706 (P).


12. Pluchea heterophylla Vatke

SOMALIA: Gedo, 14 km on road between Luuq and Beled Xaawo [c. 3°48'N, 42°33'E], 7. 6. 
1989, M. Thulin & B. A. Mohamed 6953 (K, UPS); airstrip 2 km NE of Galkayo, 6°47'N, 
47°27'E, 22.5.1979, J. B. Gillett & al. 21896 (MO); one km from Galkayo towards Garoowe, 
20.12.1977, Kazmi, Cilmi, Mahmund & Sulaiman 3 (M); Bacino del Nogal, presso la foce del 
fiume Nogal [c. 8°15'N, 49°15'E], 19.6.1924, Puccioni & Stefanini 839 [927] (FI); Region 
Nogal, 20 km NE Garoe [8°26'N, 48°30'E], 5.3.1988, B. Gabriel (B); hills SW of Las Anod 
[8°28'N, 47°21'E], 5.2.1982, M. Thulin 4203 (UPS); Region Nogal, 31 km NE Garoe, c. 500 
m, [8°30'N, 48°34'E], 5.3.1988, B. Gabriel (B); Nugal Region, Garoe-Halin, c. 8°37'N, 
48°27'E, 21.11.1985, M. Thulin & Warfa 5383 (UPS); S of Shilamadu Range [Shilemadu 
Range, Shiilowmadow: 8°59'N, 47°46'E], 38 miles E of Urygia, 28.9.1956, Bally 10870 (G); 
Burao, and within a radius of 5 km, [9°31'N, 45°33'E], 30.12.1977, Kazmi, Cilmi, Mahmund 
& Sulaiman 262 (M); 88 miles from Bosaso on Gado Rd, [c. 10°00'N, 49°10'E], 5.10.1959, C. 
F. Hemming 1638 (K); 4-15 km SW von Escushuban, [10°10'N, 50°14'E], 11/12.12.1987, B. 
Gabriel (B); Adar-Haud (Warsangeleh), 10°13'N, 48°47'E, 7.9.1929, Collenette 40 (K); 
Somaliland, 10°20'N, 45°12'E, 4.1.1933, J. B. Gillett 4805 (K); Region Bari, S-Rand des Al 
Mescat, 20 km N Escushuban, “Affenquelle”, c. 200 m, [10°22'N, 50°16'E], 23.3.1988, B. 
Gabriel (B); Karin, 10°57'N, 49°24'E, 28.10.1929, Collenette 178 (K); Sanaag, 23 km E Al 
Madjihan (Zinnmine) Richtung Karin, c. 200 m, [11°03'N, 48°56'E], 8.3.1988, B. Gabriel (B); 
Region Bari, 12 km E Bosasso [= Bender Cassim], heiße Quellen von Bio Culu, 100-200 m 
[11°14'N, 49°17'E], 5.4.1988, B. Gabriel (B).

ETHIOPIA: SIDAMO: 14 km SW of Dolo on track to Suftu, 4°08'N, 41°57'E, 19.5.1983, M. G. 
Gilbert, K. Ensermu & K. Vollesen 7607 (C, ETH); 1-10 miles W of Dolo [Dolo Odo: 4°11'N, 
42°04'E], 24.9.1953, Bally 9319 (G); Harar province, 29 miles east of Calafoh [K’elafo] on 
Calafoh-Mustahill road [c. 5°35°N, 44°20'E], 12 9.1971, J. M. Ash 1146 (ETH).


13. Pluchea indica (L.) Less

a. Pluchea indica subsp. indica

YEMEN: Hadramaut: Azzan [14°19'N, 47°27'E], 7.-8.12.1898, O. Simony (HBG); Hadramaut, 
near the old fort c. 7 km NE of Ghayl Ba Wazir, 14°49'N, 49°25'E, 21.9.1998, N. Kilian & P. 
Hein NK 5020 (B); el Lasb [? 15°12'N, 48°40'E], 5.8.1931, H. Wissmann 1431 (HBG); Terim 
– Qabr Hud [c. 16°00'N, 49°00'E], 6.8.1931, H. Wissmann 1420 (HBG); near Rahada, 
9.10.1962, G. Popov 31/1 (BM 21086).

AUSTRALIA: NORTHERN TERRITORY: Malay Bay plains [11°23'S, 132°52'E], 8.5.1987, M. J. 
Clark 1034 (MEL 713723); Gulumarri, Elcho Island, 11°56'S, 135°49'E, 18.7.1975, C. 
Dunlop 3948 (CANB 469608); Howard Island, 12°09'S, 135°26'E, 9.12.1987, C. Dunlop 
(MEL 221008); Darwin [12°27'S, 130°50'E], 12.3.1971, N. Byrnes 2087 (CANB 238290); 
South Alligator, Arnhem Hxy, 12°45'S, 132°30'E, 10.8.1983, G. Wightman 602 (CANB 
345500, 345001); Port Blane, Blue Mud Bay [13°36'S, 135°56'E], 28.1.1803, R. Brown 2099 
(BM 20937); mainland opposite Groote Eylandt [14°00'S, 136°40'E], 4.1.1803, R. Brown 
2098 (BM 20936); Arnhem Land, 13°45'S, 136°00'S, 28.11.1987, C. Dunlop 7329 (MEL 
1583141).
QUEENSLAND: Boigu Island, 9°14'S, 142°13'E, 20.10.1981, J. R. Clarkson 3876 (CANB 
345399); Andoom, Weipa, 12°38'S, 141°52'E, 19.3.1989, M. R. Okeilly 254 (MEL 119369).
PAPUA NEW GUINEA: 1.5 miles E of Delena village, Kairuku Subdistr. [8°52'S, 146°35'E], 
10.8.1962, P. J. Darbyshire 794 (A, CANB); c. 1.5 miles east of Delena village, Kairuku 
Subdistrict [8°52'S, 146°34'E], 10.8.1962, P. J. Darbyshire 794 (G); Kanosia [8°58'S, 
146°59'E], 12.1967, C. E. Carr 11011 (BM 20951, CANB); Daru Is. [9°05'S, 143°12'E], 
5.8.1971, H. Streimann 51693 (A), 27.2.1936, L. J. Brass 6216 (A); Mabadauan, Daru Distr., 
9°15'S, 142°40'E, 13.7.1968, E. E. Henty & P. Katik 38803 (A); Tupuseleia, Port Moresby, 
9°35' S, 147°20' E, 28.1.1967, A. Kairo & Streitmann NFG 35609 (A, M); Milne Bay Distict, 
Terr. of Papua [9°39'S, 149°53'E], 14.7.1954, R. D. Hoogland 4282 (A, CANB); c. 5 miles 
north of Kapa Kapa, Central District [Gabagaba: 9°48'S, 147°31'E], 8.8.1962, R. Schodde 
2731 (G); Papua, 31.5.1926, L. J. Brass 1593 (A).

VIETNAM: Saigon [10°45'N, 106°40'E], 1901, O. Debraux 215 (A); Taai Wong Mo Shan and 
vicinity Tong Fa market Ha-coi, Tonkin, [21°26'N, 107°46'E], 11-23.8.1939, W. T. Tsang 
29482 (C, LD), W. T. Tsang 29284 (A, C); Dran, South Annam [11°51'N, 108°35'E], 1918, C. 
Boden Kloss (BM 21039).

SINGAPORE: Singapore [1°17'N, 103°51'E], 28.1.1853, N. J. Anderson (S).

MALAYSIA: Sarawak, Lampangi Scheme, Santubong [1°43'N, 110°18'E], 10.4.1972, P. Chai 
31882 (A); Malacca [Melaka]: Johare [2°11'N, 102°14'E], 25.7.1919, A. Vesterdal (C), W. 
Griffith (BM 20992); Endau, Johore [2°38'N, 103°37'E], 3.3.1971, Hardial Singh 1041 (C); 
Teluk Merbau, Selangor [3°21'N, 101°15'E], 2.7.1937, C. W. Franck (C); Pulau Langkawi 
[6°19'N, 99°51'E], 12.11.1979, B. C. Stone 14257 (A); British North Borneo, Kotabelud 
[6°21'N, 116°26'E], 1910, L. S. Gibbs 4322 (BM 20985); North Borneo, Kudat [6°52'N, 
116°50'E], 19.10.1915, L. Topping 1394 (A); Perlis [c. 6°26'N, 100°12'E], 3.1910, Ridley 
14907 (BM 21100).

INDONESIA: Timor [c. 0°50'N, 123°50'E], ex Herb. Mus. Par. (C); Sumatra, Aer Djoman, 
Asahan [c. 3°00'N, 99°50'E], 7.1935, Rahmat Si Boeea 8272 (A); Sumatra, H. S. Yates 887 
(A, HBG); Luntas, Java, 1837, Hoffmansegg (G); Fort de Kock [Bukittinggi: 0°19'S, 
100°22'E], 1927, H. S. Yates 2473 (BM 21179, S); Aer Djoman, Asahan (east of Serbangan) 
[c. 3°00'N, 99°40'E], 5.8.1935, Rahmat Si Boeea 8321 (G); Insula Moluccana, Amboyna 
[Ambon: 3°43'S, 128°12'E], 1796, Ch. Smith (BM 21125); Batavia [Jakarta: 6°10'S, 
106°48'E], 15.5.1897, H. Müller (S), 28.1.1853, N. J. Anderson (S), ad Bataviam, Java, 
Mellerborg (S); Batavia, 1835, A. Delessert (G); Java, Cook’s 1th voyage (BM 21100); Java, 
Tandjong [5°13'N, 96°18'E], 16.10.1904, B. P. G. Hochreutiner 2076 (G), 12.5.1904, B. P. G. 
Hochreutiner 1128 (G); Bogor, Parung [6°25'S, 106°42'E], 1.11.1988, Harini 026 (A); H 
Bogor [6°35'S, 106°47'E], Zollinger (BR); Javae, 1893, Junghuhn 336 (A), M. Commerson 
(G); Tanimbar Is.: Pulau Selaru [c. 8°20'S, 131°00'E], 13.3.1956, D. R. Pleyte 86 (A); 
Selebes, Bare Bare, 20.2.1929, G. Kjellberg 319 (S).

THAILAND: Trip from Malacca creek to Talo Wao [c. 6°36'N, 99°37'E], G. Congdon 262 (A); 
Klong, near Trat. [6°48'N, 100°37'E], 19.1.1958, K. Larsen 539 (C); Prov. Songkhla, 
Songkhla lake [7°12'N, 100°36'E], 13.9.1984, N. Fukuoka 36111 (A); N of Phuket, 7 km [c. 
7°50'N, 98°24'E], 6.2.1966, B. Hansen & T. Smitinand 12189 (C); Phuket, hill near Thalang, 
E of road to Phuket, 8.5.1968, van Beusekom & Phengkhlai 645 (C); Phuket: Nai Harn Beach 
[c 7°53'N, 98°24'E], 14.12.1979, Shimizu, Toyokuni, Koyama & al. 27301 (C); Ko Yao Yai 
[7°52'N, 98°32'E], 21.2.1966, B. Hansen & T. Smitinand 12425 (C); Salak Pet, Kaw Chang 
[Kaw Kwang? 9°00'N, 99°56'E], 7.6.1925, Kerr 50 (BM 21026); Prachuap Khiri Khan: Ban 
Kunta Nod, Samroiyod NP [c. 11°49'N, 99°48'E], 5.12.1979, T. Shimizu & Koyama & 
Fukuoka & al. 26162 (C); S.W. Prov. Prachuap Khiri Khan: Khao Klong Wan, 23.8.1982, 
Tatemi Shimizu & al. 28748 (A); Paknam, SE of Bangkok [c. 12°29'N, 102°04'E], 7.1.1958, 
K. Larsen 83 (C); 10.3.1958, K. Larsen 2013 (C); Prachuap Kiri Khan: Sam Roy Yot, 
12°15'N, 99°55'E, 10.5.1974, K. Larsen & Supee S. Larsen 33776 (B); Samut Prakan: Ban 
Bang Pu Kao along seashore [13°36'N, 100°36'E], 14.8.1967, T. Shimizu, Koyama & Fukuoka 
7548 (A, C); Chachoengsao: Bang Pa Ken Distr. [13°42'N, 101°05'E], 30.11.1979, Shimizu, 
Toyokuni & al. 23711 (A, C); Bangkok [13°45'N, 100°31'E], Zimmermann 108 (B, BM 
21187, BR, M, S); Krungtep, Bangkok, 16.7.1966, K. Larsen 443 (C); 25.4.1919, Kerr 3674 
(BM 21030); Bangkok: Raminthra, 23.1.1983, H. Koyama & H. Terao 32872 (A); Prov. 
Pranakorn. 18 km S of Bankok, 10.8.1963, R. Merrill King 5597 (A); Nakon Chaisi [13°48'N, 
100°11'E], 6.6.1920, G. W. Groff 5819 (A); Samut Sakhon, Ban Khlong Su, 14°31'N, 
100°09'E, 26.6.1968, H. M. van der Kevie 18 (C); Siam, 2.3.1924, A. F. G. Kerr 8590 (B, BM 
21033).

BURMA: Heinze Bay [14°42'N, 97°52'E], 8.11.1924, R. N. Parker 2186 (A); Rangoon 
[16°47'N, 96°10'E], Buchanan-Hamilton (BM 20944); Jai?, 2.1901, Shaik Mokin 422 (G).

PHILIPPINES: Buayan, Cotabato Prov. Mindanao [7°18'N, 124°35'E], 4.1932, M. Ramos & 
Edano 85110 (A); Bo. Manaul, Mansalay, Oriental Mindoro [12°31'N, 121°26'E], 12.1952, 
M. D. Sulit 17148 (BM 21130, A); Zapote, Rizal Prov., Luzon [c. 14°19'N, 121°05'E], 
23.1.1947, D. Mendoza 4190 (BR), E. D. Merril 989 (A, BM 21069); Mindoro [16°55'N, 
120°25'E], 2.-4.1908, M. L. Merritt 9777 (BR), 4.1903, E. D. Merril 916 (A), 5.8.1953, H. C. 
Conklin 18625 (A).

INDIA: Calcutta [22°32'N, 88°22'E], 5.4.1874, C. B. Clarke 21646 (BM 20955), C. B. Clarke 
21639 (BM 20954); Karnal [29°41'N, 76°59'E], 2.11.1986, J. R. Drummond 25736 (E); 
Galathea-Expedition 1845-47 [?Nicobar Islands], Herb. Wallich 977 (C); Bengal, Roxburgh 
(BM 21102).

BANGLADESH: East Bengal, Herb. Griffith 3166/1 (C, GOET, M, S).

HAWAII: Kawaihafoai, Oahu, 8.4.1950, O. Degener 20935 (B); Laysan Island, 19.9.1964, C. 
R. Long 2395 (A); Islands-Kauai, 5.11.1936, E. Y. Hosaka 1662 (A); Oahu, 5.7.1969, J. 
Henrickson 3820b (A), 7.6.1963, T. S. Cooperrider 9251 (A), 4.1.1979, S. Tsugaru & Nishino 
5217 (A).

CHINA: ISL. HAINAN: Ya-xian [18°14'N, 109°30'E], 30.7.1981, T. Tuyama & al. 81314 (A); 
Hainan, Fung Leng, Ngai District, 1.9.1932, S. K. Lau 460 (A, B, BM 21042, E, G); Hainan, 
11.1889, A. Henry 3210 (G); Hainan, Hoi Hau, King-Shan Distr., [20°03'N, 110°19'E], 
2.10.1927, Tsang Wai-Tak 984, L. U. 16438 (A, E, G, S); Hainan, 23.10.1933, C. Wang 
34838 (A); Nam Shan Leng, Yai-hsien District, 7.-30.4.1935, S. K. Lau 5994 (A, E); Hainan, 
28.8.1936, S. K. Lau 27790 (A); Hainan, 11.1889, A. Henry 8708 (A); Shantou, Kwangtung 
[23°00'N, 113°00'E], 15.8.1930, T. N. Lieu L. 630 (E); Kwangtung [Guangdong], Hoi Kong 
District, in city [c. 23°00'N, 113°00'E], 4.6.1929, Tsiang Ying 2580 (G); Kwang Tung Prov. 
Canton and Vicinity [Guangzhou: 23°08'N, 113°20'E], 2.10.1917, C. O. Levine 334 (A), 
9.13.17, C. O. Levine 1276 (A, E); Canton [23°07'N, 113°15'E], 11.1885, Native Asui 328 
(BM 21103); Kwangsi [24°00'N, 109°00'E], H. Y. Liang 69911 (A); China, Ekeberg (S).
HONG KONG: Hong Kong [22°17'N, 114°09'E], 30.11.1973, Kit Yock Chan 1344 (A); 
17.12.1969, Shiu Ying Hu 9192 (A), 1853-56, C. Wright 288 (A); Un-loong [22°27'N, 
114°02'E], New Territory, 15.7.1940, Y. W. Taam 1541 (G); Hong Kong, Mai Po Marshes, 
Un-Long [22°30'N, 114°03'E], 28.10.1972, J. P. W. Woo & T. K. Woo 754 (A); Yeun Long, 
28.10.1972, J. W. Woo 196 (A); 11.10.1969, Shiu Ying Hu 8188 (A); 23.2.1972, Shiu Ying Hu 
11538 (A).
TAIWAN: Taipei CO.: Chuwei, Tanshui [25°10'N, 121°26'E], 8.7.1983, Ching-I Peng 5408 
(A); Pingtung, Cha-lo-suey [? 24°28'N, 118°25'E], 29.10.1980, K. U. Kramer, E. Zogg & H. 
Gassner 7926 (G); Formosa, 1864, R. Oldham 289 (A, BM 21075); Formosa, 15.11.1918, E. 
H. Wilson 11007 (A); Koshun, 8.9.1930, Tanaka 5485 (BM 21138).


b. Pluchea indica subsp. yemenensis King-Jones

YEMEN: Aden, Abiyan Governorate, Zingibar Guest House [12°46'N, 45°01'E], 8.2.1989, A. 
G. Miller, L. Guarino, N. Obadi, M. Hassan & N. Mohammed M8010 (E); Lahedj [Lahij: 
13°03'N, 44°53'E], 29.4.-1.5.1931, H. Wissmann 1439 (HBG); Al Huseini, north of Lahej, 
25.-30.11.1937, H. Scott & E. B. Britton 263 (BM 21116); Valleis aux environs de Lahadj, 
23.10.1889, M. Deflers 128 (BR, E, G); Wadi Bonna near Wadi Hassan 40 miles E of Aden 
on coast [c. 13°07'N, 45°27'E], 8.8.1937, A. J. Grasovsky 3 (BM 20993); Scheih O’Ahman pr. 
Aden, 6.1872, J. M. Hildebrandt 765 (BM 21003b); Abiyan Governorate: Wadi Bana 
downstream, near Yaramis [c. 13°25'N, 45°25'E], 14.3.1988, L. Boulos, A. K. Rowaished, W. 
A. Saeed & M. A. Hussein (BM 20934); hot springs 30 m[iles] E of Mudia [Mudiyah] [c. 
13°50'N, 46°10'E], 9.4.1953, A. Grierson 196 (E); Abyan, road from Lawdar to Hannap and 
Am Ayn, 14°01'N, 46°43'E, 19.1.1998, B. Mies 862 (Herb. B. Mies); Tukhana nr. Mukalla [c. 
14°30'N, 49°00'E], 1893-94, W. Lunt 64 (BM 21003a); Hadramaut: (Lassab.) el Lasb [? 
15°12'N, 48°40'E], 5.8.1931, H. Wissmann 1432 (HBG); Wadi Hajr, 100 km W of Mukalla, 
El Ain, 58 km N of Meifa Haga [14°02'N, 48°40'E], 13.2.1989, A. G. Miller, L. Guarino, N. 
Obadi, M. Hassan & N. Mohammed M8158 (E).


14. Pluchea lanceolata (DC.) C. B. Clarke

AFGHANISTAN: Prov. Kunar: Kunartal bei Asmar, [35°03'N, 71°29'E,], 5.6.1964, H. F. 
Neubauer 4545 (B); Griffith 3134 (C, GOET, L, M); Cabul [34°31'N, 69°11'E], Honigberger 
(W).

INDIA: Agra [27°11'N, 78°01'E], 3.1909, A. Meebold 11036 (W); Punjab, Karnal Jungle 
[29°41'N, 76°58'E], 1.5.1887, J. R. Drummond 25761 (E), 12.6.1886, J. R. Drummond 25764 
(E), J. R. Drummond 25759 (E); Panjab, T. Thomson (A, BM, C, E, G, LD, W); N W India, 
1871, J. L. Stewart 1662 (E); Baluchistan, Herb. George Watt 4093 (E); Cawnpore [= Kanpur 
26°28'N, 80°21'E], Madden 504 (E); Punjab, Gurgaon [Gurgaon: 28°27'N, 77°01'E], 
30.3.1887, J. R. Drummond 25760 (E); Aligarh Distr. [c. 27°54N, 78°04'E], 12.1885, J. F. 
Duthie? 4700 (a) (WU).

PAKISTAN: Clifton [24°48'N, 67°02'E], 20.4.56, S. M. H. Jafri? 1373 (E); Kinghar Lake, 
Karachi Distr., [c. 24°54'N, 68°03'E], 22.1.1934, W. Koelz 7569 (G); Jamshoro [25°26'N, 
68°20'E], 5.-6.1974, S. M. A. Kazmi 6394 (M); Larkana [27°32'N, 68°18'E], 26.4.1957, 
Ashfagali? 5302 (E); 8 miles from Rajanpur on way to D. G. Khan [29°15'N, 70°30'E], 
22.5.1972, Sultanul Abedin & al. 9606 (G); Sibi [29°33'N, 67°53'E], 3.12.1887, coll. ignot. 
4093 (E); Punjab, Multan [30°10'N, 71°33'E], 3.7.1906, coll. ignot. 25905 (E); Hasan Abdal 
[30°26'N, 72°42'E], 7.5.1925, R. R. Stewart 1157 (A); Lahore [31°34'N, 74°18'E], 9.1846, T. 
Thomson 1470 (HBG); Rawalpindi [33°36'N, 73°04'E], 4.1934, R. R. Stewart 13841 (A, MO); 
Distr. Peshawar, Michni [c. 34°11'N, 71°27'E], 29.5.1958, B. L. Burtt 695 (E), Michni 
Peshawar Road, 10.5.1966, J. Ahmad 88 (M); Spain, S. Waziristan [33°00'N, 70°00'E], 
15.7.1977, Hafizullah & Ayyaz Abbasi (A); Karachi University campus [24°52'N, 67°03'E], 
19.4.1969, Sayed Manzoor Ashray Laidi 78 (MO).

CHAD: Lac Tchad et Kanem de [N’Gourie] á Modou [c. 13°42'N, 15°19'E], 1.3/10.1903, A. 
Chevalier 10152 (B, P 11596); Lac Tchad; archipel Kivu, 7.10.1903, A. Chevalier 10144 (P 
11595); Lac Tchad de Ngouri a Madon, 1.-3.10.1903, A. Chevalier 10192 (P); Lake Chad & 
Bornu [c. 13°20'N, 14°20'E], 1.-2. 1911, P. A. Talbot 1001 (BM 21135); Gamga [12°14'N, 
15°49'E], 2.1908, Gaillard (P 11599); Moito [Moyto: 12°35'N, 16°33'E], 9.-10.9.1903, A. 
Chevalier 9812 (P 11594).

SENEGAL: Dakar, Village Almadies [14°44'N, 17°31'W], 5.11.1955, Adam Jacques-Georges 
11087 (C, MO); Gandhiolle [Gandiol: 15°50'N, 16°29'W], 1837, M. Heudelot 400 (G, P 
11582); 5.1829, Perrottet (G); 1835, Leyssieur (G); 10.2.1825, Perrottet (G); Senegal, D. 
Leprieux (P 11585).

CAPE VERDE: Ilha de S. Vicente [16°54'N, 25°00'W], 18.2.1956, Grandvaux Barbosa 6657 
(BR).


15. Pluchea linearifolia C. B. Clarke

INDIA: Lohit [Lohit River: 27°48'N, 95°28'E], Herb. Griffith (K); Assam, Abor hills, 28°15'N, 
95°30'E, 3.12.1928, F. Kingdon Ward 8752 (K); Assam, Masters (L); East Bengal, Herb. 
Griffith (C, L).


16. Pluchea lycioides (Hiern.) Merxm.

NAMIBIA: Kaokoveld, banks of Kunene 17°15'S, 12°26'E, 16.8.1956, R. Story 5825 (M)

ANGOLA: Mossamedes [15°10'S, 12°09'E], 11.1900, J. Jomreiber (K).


17. Pluchea nogalensis Chiov.

SOMALIA: Ceelbuur (El Bur), 4°40'N, 46°37'E, 25.5.1989, M. Thulin & Abdi M. Dahir 6509 
(K, UPS); Region Nogal, 31 km NE Garoe [c. 8°30'N, 48°34'E], 5.3.1988, B. Gabriel (B); 
Region Bari, 10 km W von Escushuban [10°18'N, 50°05'E], 26.3.1988, B. Gabriel (B); 
Region Bari, Strasse Garoe-Karin, bei Las Daouè im SW Darror-Tal [Laas Dawaco: 10°26'N, 
49°07'E], 6.3.1989, B. Gabriel (B); Adar-Ad (Warsangeleh), 10°33'N, 48°59'E, 16.10.1929, 
Collenette 144 (K); 33 km on the road between Hamur and Karin (Bosaso) [c. 11°10'N, 
49°10'E], 28.11.1986, M. Thulin & A. M. Warfa 6150 (K); Somaliland, 24.11.1942, Peck 342 
(K).


18. Pluchea obovata Balf.

SOCOTRA: Hajhir mountains: wadi Qishn, 12°34'N, 54°02'E, 31.3.1996, P. Hein 96-600 (B); 
Adho Demalu [Adho Dimelus: 12°35'N, 54°01'E], 16.3.1953, G. Popov 231 (BM 21091); 
Haggier Mountains, Fieri Peaks [12°36'N, 54°01'], 11.2.1990, A. G. Miller & al. M10453 (E); 
12°35'N, 54°03E, 19.4.1967, A. R. Smith & Lavranos 423 (K); Aduno pass [c. 12°35'N, 
54°05'E], 6.3.1989, A. G. Miller & al. M8666 (E, UPS); 1897, Mr. & Mrs. Theodore Bent 
(K); Wadi Daneghan, 10.2.1990, A. G. Miller & al. M10386 (UPS).


19. Pluchea odorata (L.) Cass. in Cuvier

SENEGAL: Senegambia, 1820, Perrottet (BM 21082).

AUSTRALIA: Christmas Islands: Motu Upua, 21.11.1964, C. R. Long 2753 (A).


20. Pluchea ovalis (Pers.) DC.

CAPE VERDE ISLANDS: L. Antonii [?St. Antão: 16°30'N, 23°30'W], 3.1851, J. A. Schmidt 
(HBG, S, W); Tarrafal [16°34'N, 24°22'W], 10.3.1864, R. T. Lowe (BM 21049); Ribeira 
d’Agoa, S. Nicolao [16°35'N, 24°15'W], 7.1857, C. Bolle (P 11610); Lac Retba, 5.1984, D. 
Thoen 7194 (BR).

MOROCCO: Insgane [30°21'N, 9°32'W], 6.4.1936, G. Samuelsson 6509 (S); Souss: Bords de l’ 
Oued Souss au pont des Ait Melloul [30°22'N, 9°36'W], 25.4.1936, J. Gattefossé (G, S), 
2.4.1934, R. Maire & E. Wilczek 202 (S), 12.4.1922, R. Maire 4138 (G), 10.5.1923, E. 
Jahandiez 304 (BM 21189, C, E, G, LD); Agadir [30°30'N, 9°40'W], 6.4.1936, O. Paulsen 
(C); Taroudant, [30°31'N, 8°55'W], 12.4.1922, M. Mainey ? (LD).

MAURITANIA: Toungad [20°03'N, 13°08'W], 3.5.1934, Th. Monod 304 (P 11614); Adrar de 
Mauritanie, [20°30'N, 13°30'W], 8.7.1958, A. Naegelé (B); Tergiv springs N Atar [c. 20°31'N, 
13°03'W], 5.5.59, G. Popov 58 (BM 21087).

SENEGAL: Senegambia ad paludem N’Boro [?Nianing Boro: 14°20'N, 16°55'W], 1838, 
Brunner 77 (BM 20941); Dakar [14°38'N, 17°27'E], 5.1.1928, O. Hagerup 771 (BR, C, P 
11495), 16.2.1948, A. Jacques-Georges 594 (M); Bargny, prés la Cimenterie [14°42'N, 
17°14'W], 20.5.1960, J. & A. Raynal 5878 (P11624); Dakar, nahe der Küste, 12.1935, 
Buschant (B); Rufisque [14°43'N, 17°17'W], 20.8.1951, R. P. Berhaut 352 (BR, P 11616, P 
11617); Senegambiae, Herb. Reinwardtianum (L); 1830, Leprieur (W), 1820, Perrottet (BM).

TOGO: Togo prope Lome, [6°10'N, 1°21'E], 1900/1902, Warnecke 223 (BR, G, HBG, M); 
Warnecke 312 (BR).

CAMEROON: Region de Makari, 4 km NW de Ngouma, [12°37'N, 14°18'E], 26.1.1985, S. 
Lisowski 1766 (BR); Fotokoll 100 km NW Fort Foureau, 6.10.9164, de Wilde & de Wilde-
Duyfjes 3664 (BR); Fotokoll 100 km NW Ft [Fort]Lamy, 27.11.1969, F. N. Hepper 4027 
(BR).

NIGER: Oasis 30 km d’Agadey? [c. 14°00N, 3°20'E], 8.1.1975, P. Lavie 849 (P 11639), 
Agadies, 6.2.1932, Chevalier 43477 (P 11635).

MALI: ?Gouri [13°30'N, 3°27'W], 22.7.1899, Chevalier 1204 (P 11634); Sanban [15°45'N, 
4°33'W], 1932, T. Rogeon 208 (P 11633), 9.3.1932, A. Chevalier (P 11632).

CHAD: piste Bol. N’Gouri [8°27'N, 17°17'E], 7.2.1964, Mosnier 1983 (P 11654); 90 km ONO 
de Fort-Foureau [12°05'N, 15°02'E], 6.10.1964, R. Letouzey 7223 (P 11642); N’Djamena, 
[12°10'N, 14°59'E], 23.12.1984, S. Lisowski 1644 (BR), 18.1.1985, S. Lisowski 1734 (BR); 
Miresoula (40 km WNW Massakory) [13°06'N, 15°23'E], 23.12.1964, J. & A. Raynal 12652 
(P 11655); Rombou [14°06'N, 15°14'E], 5.1.1968, J. Maley 172 (P 11649); Nord-Abdachung 
des Tarso Tousside: Mosa [c. 21°00'N, 16°30'E], 9.2.1965, H. Scholz 496 (B).

SUDAN: Hillet Nueyr, Bahr el Jebel [c. 9°00'N, 30°30'E], 2.2.1930, N. Douglas Simpson 7479 
(BM 21124); Zalingei [12°54'N, 23°29'E], 18.1.1965, de Wilde & de Wilde-Duyfjies 5336 
(BR).

ETHIOPIA: Lower Omo River Basin, [4°30'N, 36°00'E], 21.7.1969, C. J. Carr 689 (BR); Gamo 
Gofa: 24 km N of Arba Minch, 6°08'N, 37°38'E, 12.5.1986, T. Mesfin, O. Sebsebe & K. 
Vollesen 4095 (C); Lake Awasa, Sidamo, 7°00'N, 38°25'E, 10.5.1958, H. F. Mooney 7290 
(B); along Awash River, about 5km downstream from Koka Dam, 8°40'N, 39°10'E, 
29.10.1972, I. Friis, G. Gilbert, F. Rasmussen & K. Vollesen 714 (BR, C).

SOMALIA: Upper Sheikh [? 9°56'N, 45°11'E], 2.12.1919, E. Godmann 57 (BM 20987); 
Kábble, Rive del Caschei [? 0°55'N, 42°36'E], 1939, R. Corradi 2085 (W!); Neghelli, R. 
Corradi 1958 (W).

BURUNDI: Usumbura [Bujumbura: 3°22'S, 29°21'E], 16.12.1939, Verhulst 402 (BR); 
Bujumbura, 10.3.1971, J. Lewalle 5295 (BR), 28.9.1971, J. Lewalle 6143 (BR), 13.2.1972, J. 
Lewalle 6538 (BR), 30.10.1973, M. Reekmans 2862 (BR); Ruzizi, en aval de la route 
Usumbura-Uvira [3°16'S, 29°14'E], 28.1.1956, J. J. Symoens 2208 (BR), 1962, A. DeCrzene 
11342 (BR), 30.3.1975, J. Lambinon 75/57 (BR), 20.2.1980, J. Bouharmont 12932 (BR); 
Réserve naturelle de la Ruzizi, 3°16'S, 29°17'E, 25.8.1984, J. Lejoy 84/659 (BR); Bubanza, 
3°16'S, 29°18'E, 11.5.1981, M. Reekmans 11146 (BR); Bubanza, Plaine Ruzizi, 8.11.1974, M. 
Reekmans 3909 (BR); Bubanza, Gihungwe, 3.11.1974, M. Reekmans 3903 (BR); 2.1950, 
Germain 5959 (BR).

CONGO: Plaine au sud du Lac Edouard [c. 0°25'S, 29°30'E], 5-7.1929, H. Humbert 8703 bis (P 
11662); Beni to Kasindi, [Beni: 0°29'N, 29°29'E], 9.8.1914, Bequaert 5208 (BR); P.N.A. 
Plaine de la Ruindi [c. 0°47'S, 29°17'E], 28.7.1937, J. Louis 4795 (BR), 1937, J. Lebrun 7806 
(BR), Lac Tsimuka, 1.1950, R. Germain 5613 (BR); Rutshuru [1°11'S, 29°26'E], 21.2.1937, J. 
Ghesquiére 3770 (BR), 9.1937, J. Ghesquière 5310 (BR), 10.1937, J. Ghesquière 5419 (B, 
BR), Région confluent rivières Molindi et Rutshuru, 18.2.1958, H. Heine 171 (BR); Lac 
Albert, Kasenyi, [1°23'N, 30°25'E], 12.5.1953, D. van der Ben 395 (BR), 24.3.1978, S. 
Lisowski 48341 (BR), 24.3.1978, Lejoly 3152 (BR), 11.1935, Bredo 1805 (BR), Bredo 2113 
(BR), Bredo 2182 (BR); en face km 68 route Kisangani, 20.11.1982, L. Pauwels 6615 (BR).

ANGOLA: Distr. Bumbo, Serra da Chella supra Bumbo [6°55'S, 14°29'E], 10.1859, Welwitsch 
3434 (BM 21161); Bumbo, 1883, Welwitsch 3434 (BR, G).

TANZANIA: Kalehe, 2°15'S, 28°38'E, 24.5.1960, J. Petit 349 (BR); proche Sange, 2°22'S, 
28°43'E, 29.11.1955, Herb. Pierlot 995 (BR); Nassa, Mwanza [c. 2°30'S, 33°40'E], 
9.10.1952, R. Tanner 1045 (B, BR); Olduwai Headwaters [c. 2°57'S, 35°10'E], 4.12.1956, 
Greenway 9106 (B); Ngorongoro Conserv. Area: W edge of Lake Masek [c. 3°00'S, 35°30'E], 
1.10.1977, J. Raynal 19337 (BR, HBG, P 11673); Ngurdoto Nat. Park [c. 3°14'S, 36°53'E], 
5.11.1965, P. J. Greenway & Kanuri 12280 (BR); Distr. Moshi, Flu? Weru-Weru, [c. 3°20'S, 
37°18'E], 14.2.1914, A. Peter (BR); Kalundu prés d’Uviva [c. 3°26'S, 29°08'E], 12.6.1956, J. 
J. Symoens 2664 (BR); au S de Kalundu, 20.8.1955, J. J. Symoens 1272 (BR); Nassa 
Chiefdom, Mwanza District [2°24'S, 33°33'E], 21.10.1952, R. Tanner 1080 (BR); Bagayo 
River [c. 3°30'S, 35°47'E], 23.6.1965, Greenway & Kanuri 11897 (BR); Arusha-Dodoma rd. 
c. 5 km NE Magugu, 3°58'S, 35°47'E, 13.6.1990, T. Eriksson, V. Kalema & G. Leliyo TE 540 
(S); Baraka [4°07'S, 29°05'E], 17.4.1947, L. Van Meel 1629 (BR); near Lake Rukwa [c. 
7°30'S, 32°00'E], 15.8.1935, A. P. G. Michelmore 1409 (BR).

KENYA: Southern N’guaso Nyiro River, “Sotik Country”[Sotik: 0°41'S, 35°06'E], 24-
30.6.1909, E. A. Mearns 708 (BM 21067); Rift Valley, Lac Nairashe [0°46'S, 36°21'E], 1929, 
H. Humbert 9151 (P 11672); 9 km from Nairobi city on the main road to Nakuru – at the farm 
of Wellcome [1°17'S, 36°49'E], 24.6.1977, F. N. Gachathi 252 (B); Nairobi, 10.1966, 
Duvigneaud 40 Pl. 1 (BR); Athi River Township [1°26'S, 36°59'E], 24.8.1971, O. Mwangangi 
1725 (BR); Hola [1°29'S, 40°02'E], 23.10.1971, N. I. B. 25 (BR); Narok Distr. Narosura Rest 
Camp. [1°32'S, 35°52'E], 16.8.1961, Glover, Gwynne, Samuel & Tucker 2393 (BR); Bura, 
Tana River [3°30'S, 38°18'E], 5.3.1963, Njoroge Thairu 37 (BR).

RWANDA: Mohasi-See [1°51'S, 30°24'E], 7.12.1907, Mildbraed 639 (HBG); Parc National 
Akagera, Ile Ihema dans le lac Ihema [1°52'S, 30°46'E], 30.4.1973, G. Troupin 15011 (BR), 
30.5.1969, G. Bouxin & M. Radoux 497 (BR); Bugarama [2°38'S, 29°00'E], 1974, P. Auquier 
3656 (BR).

UGANDA: Toro Distr., in crater of Katwe Lake [c. 0°07'S, 29°52' E], 1.12.1934, G. Taylor 
2245 (BM 21139, S); route de M’barara a Kabale [1°15'S, 29°59'E], 10.2.1952, M. Marlier 
1221 (BR).

ZAMBIA: Kaput Distr, 8°25'S, 29°47'E, 16.4.1989, G. Pope & A. R. Smith & Goyder 2159 
(BR); Régions de Tchoma et de Mwero Wantipa, Kabwe [8°35'S, 29°34'E], 11.7.1939, Bredo 
3148 (BR); L. Chishi [8°55'S, 29°47'E], 13.9.1958, D. B. Fanshawe 4816 (BR).

NAMIBIA: Kaokoveld, Bainesberge bei Otjibemba, 17°16'S, 12°53'E, 14.7.1969, P. G. Meyer 
1303 (M), 16.6.1965, W. Giess 8978 (M).

MADAGASCAR: Vallée de L’Onilahy, vers L’Embouchure [c. 23°00'S, 44°00'E], 27.-
30.9.1924, H. Humbert 2675 (B).

COMORES: Mayotte [12°50'S, 45°10'E], Boivin 3146 (W).

SAUDI ARABIA: Qatif Oasis [26°36'N, 49°58'E], 31.3.1987, S. Collenette 6189 (E), 27.1.1966, 
J. P. Mandaville 527 (BM 21061); 2 km E of al-Hufu, al-Hasa Oasis, [25°20'N, 49°34'E], 
22.10.1987, J. P. Mandaville, Jr. 8662 (E).

OMAN: Dhofar, Central Salalah [17°01'N, 54°04'E], 17.6.1978, A. Berkeley 58 (E); Thumrait-
Hegeif T/O, Dhofar, 17°08'N, 54°08'E, 25.11.1981, J. R. Maconochie 2981 (BM 21058, E).

BAHRAIN: Between Hamad Town & the BDF hospital, [26°06'N, 50°30'E], 2.1.1985, A. M. 
Alder 86 (E); nr. oil refinery, 18.2.1984, M. Cornes 106 (E); insula Bahrein, 1.3.1893, 
Bornmüller 400 (B, BM 20930, E, HBG, LD, W).

IRAQ: Marshes of southern Iraq: Dibin, 22.4.1952, W. Thesiger 1260 (BM 21145); Al Agar 
[?Imam el Agar 32°04'N, 46°05'E], 14.5.1952, W. Thesiger 1274 (BM); Basrah, Maquil, in 
insula fluvii Shatt al-Arab [Al Ma’qil: 30°33'N, 47°49'E], 16.3.1957, K. H. Rechinger 8432 
(E, M), 4.12.1960, R. Wheeler Haines 1884 (E).

INDIA: Coimbatore [11°00'N, 76°57'E], C. E. C. Fischer 420 (B); Pondicherry [11°59'N, 
79°50'E], 6.1909, Ch. d’ Alleizette (L); Belur Maisor [c. 13°00'N, 75°50'E], 11.1908, Meebold 
9624 (E, S); Scind, Stocks (A, L, W); Belur, Staat Mysore [13°08'N, 75°51'E], 31.1.1952, 
Gilli (W); Peninsula Ind. Orientalis, Herb Wight 521 (E), 2053 (E); 1594 (C, L).


21. Pluchea pectinata F. G. Davis & J.-P. Lebrun

ETHIOPIA: Sidamo, 4°08'N, 41°57'E, 19.5.1983, M. G. Gilbert, Ensermu K., Vollesen 7607 
(K); Sidamo, 4°14'N, 42°03'E, 18.5.1983, Rippstein (K); Sidamo, ferry across the Ganale 
Daria river NE of Dolo, 4°14'N, 42°03'E, 18.5.1983, M. G. Gilbert, Ensermu, K. & K. 
Vollesen 7510 (ETH); Sidamo: 17 km from Dolo Odo to Dolo Bay near Geanle River 7 km 
off the main dolo-Negelle Road, 4°15'N, 42°03'E, 16.12.1990, Sebesebe D. & Ensermu K. 
2737 (ETH); Hararghe province, Kelafo, 5°30'N, 45°15'E, 26.8.1972, J. B. Allen & M. C. 
Hamersley 7 (ETH), 2.9.1972, J. B. Allen & M. C. Hamersley 145 (ETH); Godere [5°52'N, 
46°15'E], 6.1.1960, I.E.C.A.M.A., BH 45 (K); Hara province, 69 miles west of Gode, on road 
to Imi [c. 6°20'N, 42°30'E], 14.9.1971, Ash 1160 (ETH, K); 84 km W de Godde [c. 6°30'N, 
42°30'E], 30.11.1972, G. Rippstein (K).

SOMALIA: 3 km from Buulhawa [Buurhakaba] towards Doolo [c. 2°50'N, 44°05'E], 30.6.1978, 
Kasmi, Elmi & Rodol 799 (K, M); Hiiraan region, 4°04'N, 45°15'E, 11.10.1986, P. Kuchar 
17151 (K); Wadamago [8°52'N, 46°15'E], 4.8.1941, Peck 10 (K).


22. Pluchea pinnatifida Hook.

SOMALIA: Togga Cadow, 10°54'N, 50°54'E, 14.11.1980, J. J. Beckett 554 (K); Karim, 
10°57'N, 49°24'E, 28.10.1929, C. M. Collenette 181 (K); Region Bari, Piste Kalabeit-
Escushuban-Candala [c. 11°00'N, 49°30'E], 16.3.1989, B. Gabriel (B); Bereda [Bereeda: 
11°51'N, 51°03'E], 5.11.1959, C. F. Hemming 1801 (K).


23. Pluchea pteropoda Hemsl.

CHINA: [Hainan]: Lingshui [18°31'N, 110°01'E], 10.1935, F. C. How 73931 (A, BM 21016); 
Lam Ko Distr. and Vicinity [20°00'N, 109°30'E], 8.-26.8.1934, C. I. Lei 1388 (A); Hoi Haw 
[Haikou: 20°03'N, 110°19'E], 2.10.1927, Tsang Wai-Tak 983 (A), 18.3.1935, F. C. How 
71282 (A); Hong Kong: Mai Po Marshes [22°30'N, 114°03'E], 23.2.1972, Shiu Ying Hu 
11523 (A, K).

VIETNAM: Tourane [= Da Nang], 100 km S of Hue [16°04'N, 108°14'E], 5.-7.1927, J & M. S. 
Clemens 3645 (BM 20958, G, K).

PHILIPPINES: Bangui [18°32'N, 120°46'E], Prov. of Ilocos Norte, Luzon, 2.-3. 1917, M. Ramos 
(L).


24. Pluchea rubelliflora (F. Muell.) B. L. Rob.

AUSTRALIA: WESTERN AUSTRALIA: Carnarvon [24°51'S, 113°45'E,], J. B. Cleland (PERTH 
532738); c. 3.5 miles S of Yardie Creek H. S. [22°33'S, 113°51'E], 26.5.1965, A. S. George 
6619 (PERTH 532746); Lyndon R., Minilya-Learmonth rd. [c. 23°30'S, 114°00'E], 31.8.1960, 
A. S. George 1450 (B, PERTH 531758); Learmonth, 22°14'S, 114°06'E, 5.9.1970, A. S. 
George 10258 (PERTH 533742); Rocky Pool, Gascoyne River, 24°45'S, 114°08'E, 10.1975, 
K. F. Kenneally 4660 (CANB 290911), 15.10.1983, P. S. Short 2079 (PERTH 531901); 4 
miles N of Exmouth township [21°50'S, 114°10'E], 28.5.1965, A. S. George 6708 (PERTH 
532568); Murchison River, Galena Bridge, 27°49'S, 114°41'E, 30.9.1985, J. D’Alonzo 705 
(PERTH 1691643); Urala Station, [21°46'S, 114°49'E], 30.4.1993, V. Long WP-VL-08 
(PERTH 3174611); NW Coastal Hwy crossing Yannarie River, 22°52'S, 114°56'E, 29.9.1995, 
Hunger & Kilian 4096 (B, PERTH, etc.); 16 km W of Gascoyne Junction [25°02'S, 
115°04'E], 29.5.1969, P. G. Wilson 8431 (PERTH 531707); Banks of Ashburton R. at Globe 
Hill [22°36'S, 115°13'E], 6.10.1905, A. Morrison (A, PERTH 531928, 531987); ExSB 113, 
Pongo paddock, Towera Station, 23°10'S, 115°15'E, 6.8.1981, R. J. Cranfield 1822 (PERTH 
531715); 9 m[iles] S of Onslow [21°52'S, 115°12'E], 28.5.1962, T. E. H. Aplin 1613 (PERTH 
531944); 5 km WSW of ExSB 358, Towrana Stn., 25°35'S, 115°15'E, 25.4.1982, R. J. 
Cranfield 2104 (PERTH 522031); Barrow Island [20°46'S, 115°24'E,], 12.10.1952, F. L. Hill 
B637 (BM 21007, K), 9.1974, W. H. Butler (PERTH 531952); NW Coastal Hwy, Nanutarra 
Roadhouse, Ashburton River, 22°33'S, 115°31'E, 3.10.1995, Hunger & Kilian 4246 (B, 
PERTH, etc.); NW Coastal Hwy crossing Cane River, c. 22°00'S, 115°36'E, 4.10.1995, 
Hunger & Kilian 4250 (B, PERTH, etc.); Barlee Range, Henry River [c. 23°00'S, 116°00'E], 
17.8.1961, R. D. Royce 6530 (PERTH 531812), 15.9.1959, A. Robinson (PERTH 531839); 
Maitland R., NW Coast Highway [21°00'S, 116°30'E], 23.2.1962, A. S. George 3308 (PERTH 
531979); Fortescue Bridge, 21°29'S, 116°41'E, 19.7.1977, S. P. Pfeiffer 17 (PERTH 533858); 
7 miles turnoff to tip, Karratha, [20°44'S, 116°52'E], 10.1986, K. Glennon 284 (PERTH 
828475); Nanutarra – Wittenoom Rd crossing Beasley River, picnic area, 22°56'S, 116°57'E, 
28.9.1995, Hunger & Kilian 4095 (B, PERTH, etc.); 2 km N of Dillydadja Well, Meka 
Station, 27°26'S, 116°59'E, 21.9.1987, R. J. Cranfield 6264 (PERTH 889814); 0.5 km E of 
Mundong Mill, Berringarra Stn., 25°57'S, 117°04'E, 29.4.1986, R. J. Cranfield 5402 (PERTH 
798487); Waranoolar Pool, Harding River, 20°58'S, 117°06'E, 12.9.1981, A. S. Weston 12734 
(PERTH 532703); P. W. D. road to Harding River Dam, [c. 20°44'S, 117°09'E], 8.9.1985, P. 
Glennon 219 (PERTH 865478); Millstream [21°41'S, 117°10'E], 27.9.1969, M. I. H. Brooker 
2132 (PERTH 532681), Pool near Horseshore Gorge, Millstream-Chichester NP, 14.8.1990, 
E. Leyland (PERTH 1432060); Chichester Range NP, below Python Pool [21°20'S, 
117°14'E], 12.8.1974, J. H. Willis (CANB 341037); c. 28 km E of Roebourne, Jones River at 
Jones Bridge, NW Coastal Hwy, 20°54'S, 117°22'E, 19.9.1995, Hunger & Kilian 3740 (B, 
PERTH, etc.); Hardy River, shortly W of junction Nanutarra – Wittenoom Rd with 
Paraburdoo Rd, c. 22°55'S, 117°22'E, 28.9.1995, Hunger & Kilian 4093 (B, PERTH, etc.); E 
of Roebourne, Sherlock River, at Sherlock Bridge, NW Coastal Hwy, 20°56'S, 117°36'E, 
19.9.1995, Hunger & Kilian 3750 (B, PERTH, etc.); Depuch Island, NW of Roebourne 
[20°37'S, 117°41'E], 29.5.1962, R. D. Royce 7122 (PERTH 531790, 531847); E of 
Roebourne, Harding River, at bridge of NW Coastal Hwy, c. 20°50'S, 118°00'E, Hunger & 
Kilian 3768 (B, PERTH, etc.); Munda Ck., Mundabullangana [20°31'S, 118°04'E], 24.2.1962, 
A. S. George 3366 (PERTH 532622); road from Tom Price to Paraburdoo, first crossing of 
Bellary Creek, 28.9.1995, Hunger & Kilian 4068 (B, PERTH, etc.); Port Hedland [20°24'S, 
118°36'E], 7.1951, H. Suijdendorp 3 (PERTH 532657); 4.1905, W. V. Fitzgerald 24 (PERTH 
532665); Great Northern Hwy, c. 80 km S of Port Hedland, 21°07'S, 118°42'E, 20.9.1995, 
Hunger & Kilian 3807 (B, PERTH, etc.); Hillside Woodstock Rd to Nullagine, E of 
Woodstock Homestead, 21°38'S, 118°48'E, 21.9.1995, Hunger & Kilian 3829 (B, PERTH, 
etc.); c. 105 km (118 km by road) SSE of Port Hedland, at the Turner River [c. 21°00'S, 
118°50'E], 29.4.1977, Hj. Eichler 22538 (PERTH 3057690); Corong Creek, Woodstock Stn., 
S of Port Hedland [21°45'S, 118°50'E], 24.4.1958, N. T. Burbidge 5835 (CANB 53683, 
PERTH 531723); upper Reaches of Turner River, Woodstock Stn., 28.4.1958, N. T. Burbidge 
5920 (CANB 53682, PERTH 531936, 532649); Strelly River, [Strelly Gorge: 21°07'S, 
119°03'E], 8.-9.1907, Cleland (PERTH 531774); Hillside Woodstock Rd to Nullagine, E of 
Woodstock Homestead, 21°41'S, 119°05'E, 21.9.1995, Hunger & Kilian 3853 (B, PERTH, 
etc.); Roy Hill Rd, shortly E of crossing Port Headland – Newman Railway, c. 22°34'S, 
119°18'E, 23.9.1995, Hunger & Kilian 3942 (B, PERTH, etc.); Warralong Sta. [20°46'S, 
119°24'E], 27.5.1941, N. T. Burbidge 851 (PERTH 531782); Bonney Downs Hillside Rd to 
Nullagine crossing Shaw River, 21°48'S, 119°25'E, 21.9.1995, Hunger & Kilian 3858 (B, 
PERTH, etc.); Roy Hill Rd, 22°41'S, 119°48'E, 22.9.1995, Hunger & Kilian 3919 (B, 
PERTH, etc.); 14 m[iles] N of Marble Bar [21°00'E, 119°45'E], 9.1971, R. Mirrington 710911 
(PERTH 531995); 18 km S of Marble Bar on the Marble Bar-Pilga Road [21°30'S, 119°45'E], 
17.9.1982, N. F. Norris 890 (PERTH 532495); 21 km N on Bamboo Springs Road [22°30'S, 
120°00'E], 15.10.1978, H. Demarz 7021 (CANB 289357, K, PERTH 532584); upper waters 
of the Fortescue R., 8.5.1958, N. T. Burbidge 6047 (CANB 53684, PERTH 759155); Marble 
Bar Rd, c. 10 km S of Nullagine, 22°00'S, 120°05'E, 21.9.1995, Hunger & Kilian 3878 (B, 
PERTH, etc.); near Shay Gap (town), c. 160 km E of Port Hedland, 20°29'S, 120°09'E, 
9.7.1984, K. Newbey 10381 (MO 3658355, PERTH 531685); [near Nullagine], 21°55'S, 
120°12'E, 8.6.1979, G. Davies 80 (CANB 297084, PERTH 532541); 4 km E of Mt Hays, c. 
65 km E of Nullagine, 21°46'S, 120°45'E, 1.4.1984, K. Newbey 10110 (PERTH 531871); 10 
m[iles] W of Callawa Creek [c. 20°27'S, 120°50'E], 27.5.1947, R. D. Royce 1949 (PERTH 
531820); Yownana Creek, ca. 125 km SE of Shay Gap, 21°17'S, 121°06'E, 15.7.1984, K. 
Newbey 10498 (PERTH 532479); Oakover River, 21 km S of Two Sisters, 21°40'S, 121°08'E, 
11.7.1984, K. Newbey 10422 (CANB 352763, PERTH 531693); 29 m N of Lookout Rocks 
[21°57'S, 121°24'E], 21.5.1947, R. D. Royce 1860 (PERTH 531855); Rudall River Region, 
22°31'S, 122°05'E, 30.3.1990, R. P. Hart 906 (PERTH 1228684); Rudall River, 22°34'S, 
122°11'E, 19.5.1971, A. S. George 10669 (CANB 290077, PERTH 532533), 17.8.1971, P. G. 
Wilson 10617 (PERTH 533750); Broome, 17°57'S, 122°13'E, 27.11.1986, P. G. Wilson 12558 
(PERTH 836990); 79 km E of Broome on road to Derby, 17°49'S, 122°14'E, 13.1.1985, P. R. 
Foulkes 219 (PERTH 532460); 20 km N of Broome, and 2 km W of Beagle Bay Road on 
edge of Willie Creek, 17°46'S, 122°16'E, 12.8.1985, K. F. Kenneally 9421 (CANB 360993, 
CANB 360994, PERTH 532312); 7.5 km NW of Beagle Bay Aboriginal Community, 
16°56'S, 122°36'E, 19.8.1985, K. F. Kenneally 9445 (PERTH 532304); One Arm Point, NE 
Dampierland, 16°26'S, 123°05'E, 2.9.1987, B. J. Carter 74 (PERTH 823937); N of Dragon 
Tree Soak, Great Sandy Desert, 19°40'S, 123°21'E, 10.8.1977, A. S. George 14763 (PERTH 
532606, 532614); Derby to Broome Rd, 19.6 km S (by road) of Derby, 17°27'S, 123°44'E, 
19.4.1985, T. E. H Aplin & al. 51 (PERTH 2847272); Mt Anderson Station, Fitzroy Station 
[c. 18°01'S, 123°55'E], 7.5.1962, R. D. Royce 6905 (PERTH 532339); Augustus Is., 
Bonaparte Archipelago [15°20'S, 124°30'E], 14.5.1972, P. G. Wilson 10739 (PERTH 
532363); Kuri Bay, Bonaparte Archipelago, 15°29'S, 124°30'E, 2.9.1985, P. R. Foulkes 318 
(PERTH 864838); Coronation Island, Bonaparte Archipelago, 15°00'S, 124°55'E, 24.5.1972, 
P. G. Wilson (PERTH 532355); Careening Bay, NW Kimberley Coast, 15°07'S, 125°00'E, 
19.6.1984, K. F. Kenneally 8988 (CANB 364876, PERTH 431079); Prince Regent River 
Reserve, 15°32'S, 125°13'E, 24.8.1974, A. S. George 12612 (PERTH 759171); Hann River, 5 
miles above junction with Fitzroy River [c. 17°30'S, 125°18'E], 6.1905, W. V. Fitzgerald 1186 
(PERTH 532428); near Beverley Springs Homestead, W Kimberley, 16°43'S, 125°27'E, 
25.12.1991, R. L. Barrett 16 (PERTH 2002787); Admiralty Gulf, between Mitchell Plateau 
and Port Warrender [c. 14°30'S, 125°30'E], 24.8.1978, A. C. Beauglehole & E. G. Errey 
59084 (PERTH 829331); 3 miles S of Fitzroy Crossing [18°17'S, 125°33'E], 3.1927, A. J. F. 
(PERTH 532436); Well 33, Canning Stock Route [21°46'S, 125°39'E], 30.7.1967, A. S. 
George 9113 (PERTH 533815); Well 39, 5.5.1979, A. S. George 15613 (CANB 289945, 
PERTH 532754); “Caravan Creek” Port Warrender, Mitchell Plateau, 14°34'S, 125°50'E, 
21.6.1976, K. F. Kenneally 5190 (PERTH 492167); 180 km WNW of Halls Creek, 70 km 
NNE of Fitzroy Crossing [c. 17°30'S, 126°10'E], 25.6.1976, A. C. Beauglehole 53818 
(PERTH 830380); 6 miles NE of Mt Eliza[beth] [16°20'S, 126°20'E], 5.1905, W. V. 
Fitzgerald 140 (BM 20981, PERTH 532401); Vansittart Bay, 14°12'S, 126°22'E, 26.5.1984, 
J. H. Willis (PERTH 1691171); Sir Graham Moore Is., 13°56'S, 126°33'E, 1.7.1973, P. G. 
Wilson 11229 (PERTH 532347); Kalumburu [14°14'S, 126°38'E], 30.5.1971, N. Byrnes 2311 
(CANB 220799, K, PERTH 532320); c. 2 miles N Kalumburu Mission [14°16'S, 126°38'E], 
30.5.1971, J. R. Maconochie 1261 (CANB 217747, PERTH 532444); Drysdale River NP, +/- 
15°16'S, 126°43'E, 3.8.1975, A. S. George 13197 (PERTH 532398); WSW of Cape 
Londonderry, +/- 13°48'S, 126°46'E, 5.8.1975, A. S. George 13402 (PERTH 532371); 100 km 
W of Halls Creek, 18°43'S, 126°52'E, 8.7.1993, A. A. Mitchell 3217 (PERTH 3250148); East 
Branch Wolf Creek, +/- 60 km S of Halls Creek [c. 18°40'S, 127°38'E], 13.7.1974, G. W. 
Carr 3568 & A. C. Beauglehole 47346 (PERTH 829765); 3,5 km W Cape St. Lambert, 
14°18'S, 127°43'E, 10.6.1987, K. F. Kenneally 10312 & B. P. M. Hyland (PERTH 865958); 
Port Wyndham, Foreshore [15°30'S, 128°09'E], 9.7.1974, G. W. Carr 3273 & A. C. 
Beauglehole 47051 (PERTH 829838); Pass of the Abencerrages, Rawlinson Range, +/- 
24°50[58?]'S, 128°17'E, 4.8.1962, D. E. Symon 2479 (CANB 190021, K); 32 miles S of 
Nicholson Station [18°30'S, 128°54'E], 13.7.1949, R. A. Perry 2431 (CANB 108291); 8 km 
SE of Kununurra, 15°49'S, 128°49'E, 12.3.1978, K. Paijmans 2417 (CANB 276455).
NORTHERN TERRITORY: N of Balgo Mission, 19°43'S, 128°18'E, 20.7.1973, P. K. Latz 4041 
(PERTH 533769); The Granites [20°34'S, 130°21'E], 2.5.1958, G. Chippendale (CANB 
74631); Tanami Sanctuary, 20°52'S, 130°33'E, 25.5.1976, P. K. Latz 6475 (PERTH 531960); 
Port Darwin [= Palmerston, Darwin: 12°23'S, 130°44'E], 6.1870, Schultz 644 (K), 9.6.1886, 
T. S. Lea (BM 21045, BR), 12.1891, 1883, F. Mueller (G, WU), M. Holtze (BR); East Point, 
Darwin, 12°24'S, 130°49'E, 22.5.1979, M. O. Rankin 1848 (CANB 287710, K); 15 miles SE 
of Wavehill Station [17°33'S, 131°14'E], 17.6.1952, R. A. Perry & Lazarides 2897 (CANB 
108292, 108293, K); 6 miles SE of Mt Allan H. S. [22°18'S, 132°18'E], 24.2.1961, H. S. 
McKee 8627 (CANB 94774, K); Finke River, Palm Valley, c. 125 km SW of Alice Springs 
[24°10'S, 132°33'E], 1954, T. R. N. Lothian 180/54 (MO 1794910); Palm Valley [24°03'S, 
132°42'E], 15.4.1956, G. Chippendale (CANB 38524), 25.8.1956, G. Chippendale (CANB 
63501); Old Hamilton Downs homestead. Macdonnell Ranges, 23°38'S, 133°50'E, 
19.10.1979, D. J. Nelson 2485 (BM 21073, G); Kurundi waterhole, 20°29'S, 134°41'E, 
15.2.1972, C. R. Dunlop 2471 (CANB 239872); 1 m[ile] S Elkedra H. S. [21°11'S, 135°27'E], 
22.10.1956, G. Chippendale (CANB 38589); Gulf of Carpentaria, Rose River, 14°17'S, 
135°43'E, 7.7.1972, C. R. Dunlop 2682 (K); Underwarra [15°43'S, 136°40'E], 25.7.1967, J. 
McKean EP 58 (CANB 361102); Center Is. [15°41'S, 136°46'E], 12.10.1967, W. Braithwaite 
3288 (CANB 337592); on Barkley Hwy, 70 km NW of Barrys Caves, 20°00'S, 137°35'E, 
1.9.1981, T. & J. Whaite 3936 (K, MO 4332480).
SOUTH AUSTRALIA: Dalhousie Springs, Mt Dare fly in. [26°27'S, 135°28'E], 24.9.1974, D. E. 
Symon 9340 (B, CANB 469614); Innamincka Bore No.3, 27°44'S, 140°47'E, 28.5.1976, L. D. 
Williams 8247 (BR, CANB 351688); about 3 km E from Tuppana waterhole, 27°45'S, 
136°51'E, 29.7.1982, R. W. Purdie 2811 (CANB 331437); Neals river, 56 km S of 
Oodnadatta, [28°00'S, 135°27'E], B. Nordenstam & A. Anderberg 969 (MEL 1593636).
NEW SOUTH WALES: Pinaroo Lake = Fort Grey Basin, 29°06'S, 141°14'E, 8.5.1977, E. N. S. 
Jackson 2855 (G).
QUEENSLAND: Bowen [20°00'E, 148°10'E], 16.1.1931, C. E. Hubbard & C. W. Winders 6564 
(K); Broad Sound [22°10'S, 149°45'E], 15.9.1802, R. Brown 2102 (BM 20939[a], K); 
Carpentaria, Island [Sweer’s Island: 17°05'S, 139°35'E], 27.11.1802, R. Brown (BM 
20939[b]); near Mt Faraday [24°54'S, 147°18'E], 10.1846, T. L. Mitchell (BM 21070 [b]).

Specimens of putative hybrids between P. rubelliflora and P. dentex examined
NORTHERN TERRITORY: Muggs Mistake, Macarthur River, 15°51'S, 136°37'E, 4.7.1984, G. 
Wightman 1543 (CANB 353012); Maggie Springs, Ayer's Rock [25°21'S, 131°02'E], 
25.11.1954, G. Chippendale (CANB 33370), 16.6.1989, J. Palmer 220 (CANB 387610).
WESTERN AUSTRALIA: Burrup Peninsula, c. 20°33'S, 116°50'E, 6.10.1995, Hunger & Kilian 
4285 (B, PERTH, etc.); NW Coastal Hwy crossing Warramboo Creek, c. 21°40'S, 115°47'E, 
4.10.1995, Hunger & Kilian 4253 (B, PERTH, etc.); 5 miles E of Nullagine [21°55'S, 
120°12'E], 30.8.1966, J. V. Blockley 388 (CANB 469609); Hammersley range. 7 km on 
Wittenoom gorge road [c. 22°20'S, 118°20'E], 5.10.1989, B. Nordenstam & A. Anderberg 325 
(PERTH 1934112); Karijini NP, Kalamina Gorge, c. 22°23'S, 118°24'E, 26.9.1995, Hunger & 
Kilian 4035 (B, PERTH, etc.); Tom Price Railway Rd, 26 km NW of  Tom Price, 22°35'S, 
117°40'E, 25.9.1995, Hunger & Kilian 4033 (B, PERTH, etc.).


25. Pluchea rufescens (DC.) A. J. Scott

MADAGASCAR: Vallée De L’Antsahabe, affluent de la Lok Oho [c. 14°20'S, 49°30'E], 10.-
11.3.1949, Humbert 23316 (P); Manahor [?Manahara: Manara 15°13'S, 47°12'E], Humblot 
225 (K, P); sur la Maningory á l’entrée du lac Alaotra [17°20'S, 48°38'E], 4.1912, H. Perrier 
de la Bathie 2835 (P); Ambatondrazaka [17°49'S, 48°28'E], 18.5.1955, Botoalina 7579-RN 
(P); Ivoloina, District: Tamatave [18°05'S, 49°46'E], 11.1926, Waterlot 1004 (P); Mandera á 
Antsabe, environs de Fort-Dauphin [25°01'S, 47°00'E], 4.4.1949, M. G. Cours 3603 (P); 
Madagascar, Chapelier (P).


26. Pluchea sordida (Vatke) Oliv. & Hiern

YEMEN: Wadi Hadhraan near Taiz [c. 13°34'N, 44°02'E], 26.12.1975, J. R. I Wood Y/75/1005 
(BM 21176).

ETHIOPIA: Arussi, Shoa province, 7°35'N, 38°45'E, 8.12.1974, J. Ash 2774 (K, MO); Sodere, 
8°25'N, 39°24'E, 23.11.1975, J. W. Ash 3335 (MO).

CONGO: Base de Montagnes au SW du Lac Edouardo, Kivu, [c. 0°40'S, 29°30'E], 1929, 
Humbert 8750 (B, BR, P 11515); May ya Moto [Mai ya Moto: 0°54'S, 29°21'E], 9.1937, J. 
Lebrun 7633 (B, BR, BM 21142, C, K, MO), 26.2.1957, de Witte 14195 (BR); environs de 
Rutshuru [Rutshuru: 1°11'S, 29°28'E], 1.1932, J. Lebrun 4865 (BR); Kivu, Parc National des 
Virunga, [c. 1°30'S, 29°10'E], 16.4.1973, S. Lisowski 18071 (BR); marais de Luvungi [2°51'S, 
28°58'E], 28.2.1960, F. L. Hendrickx 7641 (BR, C); Plaine de la Ruzizi, lac Tsimuka, [c. 
3°00'S, 29°10'E], 1.1950, R. Germain 5586 (BR, K).

UGANDA: Duamsikizi Crater Lake, Toro [0°05'S, 29°59'E], 8.1941, W. J. Eggeling 4521 (K); 
Bwamba hot springs [c. 0°50'N, 30°06'E], 21.12.1969, K. Arnstein Lye 4736 (K); Katwe, 
Ruwenzori region [0°80'S, 29°53'E], 11.12.1925, T. D. Maitland 993 (BR, K).

KENYA: Shores of lake Nakuru [c. 0°21'S, 36°05'E], A. Baraza (MO), 13.4.1975, J. Lambino 
75/248 (BR); S end Lake Elmenteila [c. 0°26'S, 36°15'E], 28.10.1972, T. B. Gillet 20055 (BR, 
K); Hunter’s Lodge [2°12'S, 37°43'E], 2.5.1991, Freidberg & Kaplan 31 (K); Kitau, Manda 
Island [c. 2°17'S, 40°57'E], 12.2.1956, P. J. Grennway & S. P. Rawlins 8900 (K); pools about 
2 km S of Ngao [c. 2°24'S, 40°12'E], 1.3.1977, S. S. Hooper & Townsend 1119 (C); Amboseli 
Reserve, at Ol Tukai [2°40'S, 37°17'E], 14.9.1954, Bally 9861 (G); Tsavo East NP [c. 3°00'S, 
38°45'E], 2.9.1976, F. N. Gachathi 76/199 (B); Magarini, 3°03'S, 40°02'E, 23.9.1982, S. A. 
Robertson 3412 (K); Kilifi Dist. [c. 3°10'S, 39°40'E], 12.1975, P. Bally 16952 (C, K), 
10.10.1945, Jeffery, G. M. (PRE); Buchuma Res. Station [3°38'S, 38°54'E], 7.5.1971, S. K. 
Nganga 129 (K); Mowesa [Mwawesa: 3°57'S, 39°37'E], R. M. Graham 1788 (K, MO); 
Kinango, S of Kwale [4°08'S, 39°19'E], 30.3.1973, G. W. Sangai EA 15803 (M, MO); Kitoni 
[4°27'S, 39°19'E], 7.1967, J. Makin 401 (K).

BURUNDI: Plaine Ruzizi, Bubanza, marais Kajeke [Bubanza: 3°05'S, 29°22'E], 8.11.1974, M. 
Reekmans 3913 (BR, MO), 25.11.1974, M. Reekmans 3981 (MO), 18.5.1975, M. Reekmans 
4577 (BR, K, MO), 29.3.1975, J. Lambinon 75/43 (BR, MO); Gihanga, 3°15'S, 29°20'E, 
6.3.1976, M. Reekmans 4830 (BR, MO); Prov. Bujumbura, Katumba [Burundi], 3°20'S, 
29°18'E, 3.1979, M. Reekmans 7696 (BR, MO); Bujumbura, Katumba, 3°22'S, 29°16'E, 
5.3.1977, M. Reekmans 5785 (BR, C, MO); Plaine de la Ruzizi, Bujumbura, [3°22'S, 
29°19'E], 5.4.1966, J. Lewalle 712 (BR), 28.9.1966, J. Lewalle 1059 (BR, K, M); Yatumba, 
Burundi, Uvira [3°24'S, 29°20'E], 1.1.1991, J. de Laet H26 (BR).

TANZANIA: Masailand, near Mt Longido [2°41'S, 36°42'E], 18.12.1968, M. Richards 23372 
(K); Mbarika Mwanza [2°55'S, 32°51'E], 5.11.1952, R. Tanner 1132 (K); W of Soito 
Mudison, Serengeti Plains [c. 2°56'S, 34°54'E], 21.11.1956, P. J. Greenway 19045 (B); 
Shinyanga Distr., Ngorongoro Conserv. area [c. 3°00'S, 35°30'E], 1.10.1977, J. Raynal 19344 
(BR, HBG, K, P 11518); Ngare Nanyuki road [3°09'S, 36°51'E], 7.12.1966, H. M. Richards 
21658 (K); Lekuruki area [3°16'S, 36°58'E], 27.3.1971, H. M. Richards & Arasululu 26864 
(K); Arusha Chini, 12 miles S of Moshi [3°35' S, 37°21' E], 22.10.1964, J. Beesley 38 (M); 
Maji Moto, Lake Manyara Nat. Park [3°40'S, 35°44'E], 8.11.1963, Greenway & Kirrika 
10984 (K, M); Lushoto Distr., Lake Manka [4°44'S, 38°05'E], 12.1.1967, M. Richards 21960 
(K, M); Moa, coast, [4°46'S, 39°10'E], 12.3.1948, Bally 5796 (G); Kitolo, 14 miles N of 
Nguruka, 8.10.1949, P. R. O. Bally 7560 (K); 3 miles of Mombo, [Mombo: 4°53'S, 38°17'E], 
29.4.1953, R. B. Drummond & J. H. Hemsley 2290 (B, BR), 26.7.1970, H. Faulkner 4432 (B, 
MO); Amboni, [5°03'S, 39°03'E], 6.1893, C. Holst 2747 (K, M, W); Usambara, Lutindi 
[5°04'S, 38°22'E], 7.1893, C. Holst 3489 (BM 21010, G, JE, PRE, S); Tanga [5°04'S, 
39°06'E], 12.1913, C. F. M. Swynnerton (BM 21134); Sawa Luk, [5°07'S, 39°06'E], 
30.5.1956, Faulkner 1876 (B, BR, S); am Pangani bei Korogwe, [Korogwe: 5°09'S, 38°28'E], 
21.5.1914, A. Peter (BR), 25.2.1916, A. Peter (BR), 26.8.1976, M. E. Archbold 2163 (BR, 
MO), 13.8.1971, Magogo 116 (BR); T3 Korogwe-Handeni road, 7 km, alt. 350 m, [5°15'S, 
38°30'E,], 26.8.1976, M. E. Archbold 2163 (B); Segera [5°18'S, 38°33'E], 16.8.1971, M. E. 
Archbold 1438 (MO); Kigombe [5°18'S, 39°01'E], 13.10.1957, H. G. Faulkner 2075 (B, BR, 
K); Pangani, [Pangani Falls: 5°21'S, 38°39'E], 1.83, Stuhlmann (HBG); Muhesa, Usambara 
[5°26'S, 38°59'E], 10.4.1903, W. Busse 2143 (BR), 1.3.1943, A. Lawabeé (BR); Mswaki 
Handeni Distr., 5°28'S, 37°46'E, 15.7.1960, Sembei, J. R. 3035 (PRE); Ushongo, Mwera 
[5°31'S, 38°58'E], 7.9.1955, R. Tanner 2135 (BR, K); one mile W of Lumi River on the 
Himo-Tonga Rd. [c. 6°09'S, 37°35'E], 15.4.1958, P. Greenway 9534 (BR, K, MO); Kunduchi 
[6°40'S, 39°13'E], 3.3.1971, Wingfield 1195 (MO); Kawe – Msasani near D’Salaam, [6°45'S, 
39°17' E], 31.8.1968, B. J. Harris 2176 (C); Mchungu forest, 7°40'S, 39°17'E, 15.8.1990, 
Coastal Research Programme 1284 (MO); Mchangani, Mafia Island, 7°50'S, 39°50'E, 
20.9.1937, Greenway 5279 (PRE); Bezirk Rufiji: Rufiji oberhalb des Deltas [c. 8°00'S, 
38°45'E], 30.6.1932, H. J. Schlieben 2543 (B, BR, MO); T8 Kingupira Forest, 8°30'S, 
38°34'E, 16.6.1975, K. Vollesen 2442 (C).

ZANZIBAR: Massazini, [6° 12'S, 39° 12'E], 5.6.1960, H. G. Faulkner 2586 (B, BR, K, S); 
Zanzibar, R. P. Sagleux (P 11680, 11682, 11684).


27. Pluchea succulenta Mesfin

SOMALIA: Nugaal region, 3 km N of Bedey, E of Eil, 8°3'N, 49°57'E, 4.10.1985, Lavranos & 
Carter 23503 (K).


28. Pluchea tetranthera F. Muell.

AUSTRALIA: Western Australia: Mallina Yandeyarra road, near Mallina Station, 20°52'S, 
118°05'E, 20.9.1995, Hunger & Kilian 3780 (B, PERTH, etc.); Yale River West Picnic Area, 
20°43'S, 118°16'E, 20.9.1995, Hunger & Kilian 3786 (B, PERTH, etc.); North West Coastal 
Hyw, c. 3 km E of Yale River West Bridge, 20°40'S, 118°19'E, 20.9.1995, Hunger & Kilian 
3794 (B, PERTH, etc.); Great Northern Hyw, c. 38 km S from junction with North West 
Coastal Hwy, 20°54'S, 118°35'E, 20.9.1995, Hunger & Kilian 3797 (B, PERTH, etc.); 18 km 
N of Port Hedland [20°20'S, 118°36'E], 7.10.1985, H. Demarz 10870 (PERTH 887692); near 
Marble Bar [21°11'S, 119°44'E], 11.9.1968, J. V. Blockley 953 (PERTH 533793); 10 miles N 
of Nullagine [21°45'S, 120°06'E], 26.10.1973, H. Demarz 4752 (PERTH 759244); 10 km NE 
of Nita Downs, 19°01'S, 121°45'E, 8.10.1984, P. R. Foulkes 15 (PERTH 533130); W 
Dampier Peninsula, 17°40'S, 122°12'E, 8.10.1984, P. R. Foulkes 14 (PERTH 533122); SE of 
Broome, 18°28'S, 123°03'E, 17.8.1976, K. F. Kenneally 5709 (CANB 289673, PERTH 
533106); R2 Edgar Range Site, SE of Broome, 18°49'S, 123°17'E, 7.8.1976, K. F. Kenneally 
5525 (CANB 289723, K, PERTH 533092); McLarty Hills, 19°30'S, 123°30'E, 6.8.1977, A. S. 
George 14671 (PERTH 533645); 16 miles N of Noonkanbah Station [18°33'S, 124°51'E], 
13.9.1959, M. Lazarides 6529 (AD 96120021, CANB 85547, K, PERTH 759252); 76 miles 
W of Margaret River, SSE of Fitzroy Crossing, [c. 18°13'S, 125°33'E], 1.8.1965, A. C. 
Beanglehole 11150 (PERTH 533076); Gogo Station [18°17'S, 125°35'E], 5.10.1952, K. 
Fitzgerald (PERTH 532835); 81-97 km NW of Tanami, 19°02'S, 128°05'E, 8.1971, C. H. 
Gittins 2398 (K).
NORTHERN TERRITORY: 114,2 km WNW Tanami [19°20'S, 129°00'E], 13.9.1971, C. R. 
Dunlop 2337 (CANB 220520); South of Mongrel Downs Station, 20°56'S, 129°24'E, 
5.8.1976, P. K. Latz 6565 (AD 97733017).


29. Pluchea wallichiana DC.

INDIA: India orientalis, Wallich (BM 21158); India, 1819-20, Wallich (BM 21157); Punjab [c. 
30°00'N, 74°00'E], 18.4.1904, J. R. Drummond (G).





Species of uncertain status:

Pluchea incisa Elmer

PHILIPPINES: Baguio, Prov. Benguet, 3.1907, A. D. E. Elmer 8786 (G).


Pluchea polygonata (DC.) Gagnep.

Selected specimens examined
THAILAND: Sisawat, Kanburi [14°01'N, 99°32'E], 15.1.1926, Kerr 10215 (BM 21034); 
Chaiyphum [15°46'N, 101°55'E], 30.1.1931, Kerr 19968 (BM 21035); Huf Taleng, Korat 
[=Nakhon Ratchasima: 15°00'N, 102°06'E], 21.12.1928, Herb. Kerr 2180 (BM 21029); 
Buriram [15°00'N, 103°07'E], 26.11.1976, C. Phengklai & al. 3435 (A); between Nam Pheom 
and Tunkamang, 16°20'N, 101°45'E, 13.12.1971, C. F. van Beusekom, C. Phengkhlai, R. 
Geesink & B. Wongwan 4202 (C, K, L, MO); Sakonnakhon, 17°00'N, 104°00'E, 13.11.1984, 
G. Murata & al. 49815 (A); Djeng-Mai [Chiang-Mai: 18°47'N, 98°59'E]; am Doi-Sutep, 
28.12.1904, C. C. Housseus 290 (BM 21014, C, K, MO, S); Doi Sootep, 1.2.1910, A. F. G. 
Kerr 916 (BM 21027); Chiang Mai, 12.1957, C. G. G. J. van Steenis 19620 (L); Northern 
Chiang Mai, Doi Inthanond, 18.12.1964, K. Bunchuai 1464 (C, L); Payap. Chiangmai, 
16.12.1965, M. Tagawa, Iwatsuki & Fukuoka 2311 (A); Cangwat Chaing Mai, Amphoe 
Muang, 19.10.1978, Bojrnland & Schumacher 402 (C); Doi Chiang Dao NE, foothills at Sop 
Huay Pa Dang-Huay Na Lao Forest Station [c. 19°23'N, 98°54'E], 25.11.1996, J. F. Maxwell 
89-1443 (A, MO); N Prov. Mae Hong Son: Pai Wildelife Sanctuary Station [c. 20°00'N, 
100°50'E], 16.1.1983, H. Koyama & al. 32604 (A); Na Moutai, Prov. Petiamloth ?, 
20.12.1905, C. C. Housseus 725 (BM 21015).

BURMA: Karen Country and Hills [c. 18°00'N, 97°00'E], S. Kurz 861 (BM 21040, L); Prome, 
Htone ye Res [c. 18°45'N, 95°15'E], 7.1.1971, U. Mg. Gale 12248 (L).

LAOS: Xieng Kouang [Xieng Khouang: 19°21'N, 103°23'E], 12.1903, C. J. Spire 1534 (P); 
bassin du Se-Moun (Laos meridional), 1876, Harmand 337 (A); Pak-moun, Me-Kong, 
1866/68, C. Thorel 2737 (K, P); bassin du Se-Moun, Nord de Compong “Loai” ou “Xoai” 
[Ban Xoai?: 16°45'N, 106°23'E], 1875/77, F. J. Harmand 337 (P); Laos, 23.12.1890, M. 
Massie (BM 21066, P); Whuang ?rubang, 1890, L. Pierre (P).






Species included in the phylogenetic analysis:

Karelinia caspia (Pall.) Less.

Selected specimens examined
IRAN: Prov. Khorasan [c. 34°30'N, 58°00'E], 25./27.7.1937, K. H. Rechinger 1813 (B), 
20.8.1885, J. E. T. Aitchison 786 (C); Gorgan Province, Shore of brackish laks at Tangeli 
[37°24'N, 54°37'E], 13.10.1977, T. F. Hewer 4052 (G).

UZBEKISTAN: Reg. Samarkand, Dominium Buchara, prov. Kurban-tube [c. 39°40'N, 66°57'E], 
14.8.1913, J. Bornmüller 1032 (B); in the zapovednik Vardanze, near the village Shafrikan 
[40°07'N, 64°30'E], 29.7.1985, T. S. Elias, Murray & Newcombe 9996 (C).

MONGOLIA: Gobi, 1886, G. N. Potanin (E, G, W).

TURKMENISTAN: Bakharden, 16 km ad orientem versus a vico Bami [38°44'N, 56°48'E], 
16.9.1976, V. Vasák (W).


Streptoglossa bubakii (Domin) Dunlop

Selected specimens examined
AUSTRALIA: Pilbara, Rd to Nullagine, 2-5 km from junction with Great Northern Hwy, c. 
21°07'S, 118°42'E, 21.9.1995, Hunger & Kilian 3826 (B, PERTH); Pilbara, Tom Price 
Railway Rd, 26 km NW of Tom Price. Creek shortly N of junction with Nanutarra – 
Wittenoom Rd, 25.9.1995, Hunger & Kilian 4032 (B, PERTH).