Selected additional specimens seen


Astragalus sect. Adiaspastus

1. Astragalus albispinus Širj. & Bornm.
1a. Astragalus albispinus subsp. albispinus 

IRAN: PROV. ESFAHAN: Najafabad, Tiran to Shahr Kord, Mosiabad area, 2200 m, 27.7.1993, Maassoumi & al. 71913 (MSB); 
Ghohrud, on the road to Maravand, 2670 m, 4.9.1993, Maassoumi & Zarre 72016 (MSB, TARI). — PROV. BAKHTIARI: 
Goushaki to Gandoman, 5.6.1973, Iranshahr & Moussavi 15549 (W); 36-40 km on the road from Mobarakeh to Boroujen, 
Gardaneh-ye Dozdan, 2100-2400 m, 3.9.1993, Maassoumi & Zarre 71999 and 72006 (MSB, TARI); S de Shahr Kord, 2050 
m, 8.7.1959, Pabot 2239 (G); Shalamzar, S Shahr Kord, 2050 m, 8.7.1959, Pabot 2240 (G); a Sefid Dasht 12 km versus 
Shahr Kord, 2.6.1974, Rechinger 47148 (M, W); 10-20 km from Shahr Kord to Zarrin Shahr, 2000-2250 m, 29.8.1993, Zarre 
72959 (MSB, TARI, TUH); Farsan to Chehelgerd, 2 km after Filabad, 45 km to Chehelgerd, 2000 m, 17.7.1994, Zarre 17769 
(MSB, TARI, TUH). — PROV. YAZD: Road of Taft, Deh-e Bala, Pachenar, 2650 m, 2.9.1993, Maassoumi & Zarre 71975 
(MSB, TARI); ibid., Hedesh, 2270 m, Maassoumi & Zarre 71983 and 71984 (MSB, TARI). — PROV. FARS: Shirkuh, 20 km 
SSW of Taft, pass between Taft and Shekh Ali Shah, 2300 m, 25.5.1977, Aryavand & al. 1377 (E); 63 km to Abadeh from 
Shiraz, 2150 m, 14.7.1994, Chehregani & Zarre 17901 (MSB, TARI, TUH); Pasargad, 15 km N of Saadatabad, 2000 m, 
21.5.1964, Grant 128 (herb. Shiraz Univ., W); Eghlid, Namadan plain, Bazubacheh, 2250 m, 10.3.1921, Hatami 723 (herb. 
Shiraz Univ.); Dasht-e Arjan, NW von Shiraz, 1978, Racheli (MSB).
NOT TRACED: Persia australis, Aucher-Eloy 1279 (K: mixed with A. floccosus); Felsensteppe bei Assupas, 1.9.1886, Stapf (K, 
W: sub A. pyri nom. nud.).


1b. Astragalus albispinus subsp. esfandiarii (Širj. & Rech.f.) Zarre
Only known from the type gathering.


2. Astragalus asaphes Bunge

UZBEKISTAN: Iter Angrenense, 1909, Borodin & Kallistow (LE); iter ad distr. Namangan, 29.6.1912, Knorring 431 (LE); 
Chatkalbskii khrebet, 24.6.1939, Kudrjashev 274 (LE); Kuraminskii khrebet, 10.7.1939, Kudrjashev 591 (LE); distr. 
Taschkent, 8.7.1914, Z. von Minkwitz 1492 (LE); Mazar bei Tashkent, Regel (LE); Tian-Schan occid. jugum Kuraminskyi ad 
trajectum Kambschik in lapidosis, 2300 m, 2.6.1965, Skvortsov (M); ibid., 2300-2800 m, 28.7.1973, Vašák (G); Tashkent, 
upper part of Sur Darya, Arasan, 18.7.1909, Worodin & Kalisrov 174 (MSB).
TADZHIKISTAN: Kuraminskyi Khrebet, inter Kamguk et Ferganeskou, 28.6.1959, Botzansev 575 (LE); ibid., Knorring 431 
(LE); Daschti Kaze, 13.6.1881, Capus (P); Turkestan, Ouranmitane, 15.6.1881, Capus 351 (P: photo MSB).


3. Astragalus aureus Willd.

RUSSIA: CHECHENO-INGUSHETIA: Terek, 21.6.1893, Lipsky (LE). — DAGESTAN: Distr. Kürinskii, inter pagos Keredsh et 
Ussuck, 6300', 23.7.1898, Alexeenko (LE); distr. Samur, in decliv. jugi Buga-dag inter pagum Fija et Maza, 7800', 19.8.1900, 
Alexeenko 9319 (LE); Kurush, Becker (G-BOIS, M, P, WU); inter Rutuiloig et Chilazo, 27.6.1911, Kusnezow & Popov (LE); 
Rutulskyi region, near village Khlud, between villages Aghty and Rutulan, right bank of river Samus, 16.6.1966, Tsvelev & 
al. (LE).
GRUZIA: In collibus aridis arenosis ditionis Swant, 4000-6000', 1836, Hohenacker (G-BOIS, WU); Elbrus mts, beginning of 
the canyon Asul-su, 12.8.1957, Kerpechnikov (LE).
ARMENIA: Alagez [Aragats], Nnzhnego Kosha-bulaga, 2100 m, 19.8.1932, E. A. & N. A. Bush (LE); Megrinskii raion, Dolina 
pr. Wagrawar-tschai-prawni pritok, Otkritaja Bershina khrebet, 1.7.1956, Egorova & al. 1615 (LE); Megrinskii raion, 
Zangezurskii khrebet, 6.6.1953, Egorova & al. 1999 (LE); Abovjanskyi raion, Gegamskyi khrebet, alto planitie 10 km viarii 
ENE supra pag. Artis versus montem Azdaak, c. 14 km viarii NE supra p. Goht, 2550 m, 28.7.1981, M. A. Fischer 532 
(MSB, WU); in latere meridionali montis Aragac secus viam ad lacum Kari ducentem, 2900 m, 13.7.1975, Gabrielian 12829 
(G); distr. Zangezur, inter Litshkvas et Migry, 31.5.1923, Grossheim (LE); distr. Migri, inter m. Ketshmas et p. Tashtyn, 
2500 m, 20.8.1932, Karjagin (LE); distr. Migri, pag. Migri ad ripam fl. Migri-tschai, 29.5.1934, Karjagin (LE); Erevan via 
Davis, 5.7.1970, Ledingham 5241 (LE); Dyman, pr. Baku, 20.5.1898, Lewandowsky (LE); pro. Sevan, prope Elenovka, m. 
Bugda-tapa, 7894', 5.7.1927, Schelkovnikov & Kara-Murza (LE); on shortly turfer hillside, Aragats mts, above Antarut, c. 
1920 m, 22.6.1985, Townsend 85/780 (K); distr. Sevan, ad orientem versus pagi Covagjukh, 2000-2300 m, 5.7.1982, Vašák 
(G); distr. Ararat, montes Gegamski khrebet, in vicinitate ruinarum pagi Akhkeng, 1800-2100 m, 10.7.1975, Vašák (B, G, 
BR, M); Montes Pambakski khrebet, vicinitate oppidi Sevan, 1900-2200 m, 17.7.1975, Vašák (M); distr. Abovyan, montes 
Gegamski khrebet, 10 km ad boreo-orientem pagi Gokht versus, 1800-2000 m, 19.7.1975, Vašák (B, G, M); in declivibus 
montis Aragac, apud ruinas Amberd, 2100-2300 m, 23.7.1975, Vašák (BR, M); distr. Vardensis (Basargechar), montes 
Vardenskii khrebet, in declivibus montis Kungukh-tar, apud rupem Shish-pal., 2100-2200 m, 29.7.1975, Vašák (B, BR, G, 
M); distr. Krasnoselskoie, montes Aregani, in vicinitate pagi Tokludza, 2000-2300 m, 31.7.1975, Vašák (B, BR, G); distr. 
Nor-Bajazet, in jugo inter pagos Janych et Karawan-Saray, 28.7.1929, Zedelmeier & Heidemann (LE).
AZERBAIJAN: Gub. Baku distr. Geokszai, in valle supra p. Dschuljan infer ad m. Küpücz versus, 6500', 2.8.1890, Alexeenko 
3294 (LE); Gub. Baku, Talysch, prope pagum Orant, ad catarractum Pjolo, 5700', 5.8.1897, Alexeenko (LE); Gub. Baku, 
Talysch ad fines persicus, inter Karabagh-jurt et Mara-jurt, 7900-8200', 7.8.1897, Alexeenko (LE); Nakhichevan, Batabat, pr. 
Farkhadevi, 16.7.1969, Babajev (LE); Baku, Becker (BR, G-BOIS); prov. Erivan and Elisabethpol, circa loac. Goktscha, 
8.1897, Chociatowski (LE); Schachboz, 20.9.1949, Gadiev (LE); distr. Nachitchevan, in jugo Zangezur, in mte Salavarty, 
3000 m, 12.8.1927, Gavilov & Doluchanov (LE); Karabach, m. Ziarat, c. 2000 m, 4.7.1932, Karjagin & Mikubin (LE); 
Karabach, in mte Kyrs, 14.7.1929, Kolakovsky (LE); Talish, Meyer 252 (M, P, W); Kusarski region, the valley N river 
Kusartchai, SE slope of mt. Shachdag, 16.7.1978, Popova & Menitskii (LE); prov. Baku, distr. Lenkoran, 16.7.1930, Prilipko 
(LE); prov. Baku, distr. Kuba in m. Shach-dagh inter pass Yachja et Kurush-Eilag, 25.8.1829, Sachokjia (LE); in saxosis 
montanis et in campestribus Ghierus et fl. Mazartchai in prov. Karabagh, Szovits 255 (E, G-BOIS, L, LE, M, P, W).
TURKEY: B8 ERZURUM: Palandöcken Dagi unmittelbar S der Stadt Erzurum, Ruine gegenüber dem Gipfel mit der 
Antennenstation, c. 2900 m, Ehrendorfer & al. 787-114-36b-c (WU); Tortum-Oltu, 1 km E Kireçi geçidi, 2220 m, 15.7.1989, 
Nydegger 44440 (MSB); in montibus Tech-Dagh supra Erzeroum, 7000-8000', 7.1853, Huet Du Pavillon (G-DC, LE, P); 
Palandökendaglari geçidi N Aziziye, 2855 m, 19.7.1996, Rasmont 103 (LG); NE Erzurum, Kargapazari, NE Senyurt, 2750 
m, 20.7.1996, Rasmont 120 (LG); Turetskaj Armenia, Erzerumskii sandzhak, perewal Tashligeduk, 11.7.1916, Sapozhnikov 
& Schischkin (LE); Erzeroum, 1854, Tchihatchef 737 (G-DC); Karakaya Dag, 15 km N of Hinis, 2200 m, 19.6.1977, Watson 
5649 (E). — A9 KARS: Mts E of Kagizman, N side of pass between Akchay and Cumachy, 2350 m, 17.7.1966, Davis 46765 
(E, M); Sarikamis-Karaurgan, 10 km W Sarikamis, 6.7.1992, 2240 m, Nydegger 46622 (MSB). — B8 BINGÖL: W side of 
Bingöl Dag, above Karliova, 2500 m, 10.7.1966, Davis 46183 (E, M). — A9 AGRI: Agri-Horasan, W Eleskirt, Weg nach 
Hayrangöl, 2450 m, 17.8.1987, Engel (MSB). — B9 BITLIS: Adilcevaz, Harmanteppe village, Süphan Dagi, c. 3000 m, 
7.8.1989, Aytaç & Metzger 2979 (GAZI); Süphan Dag, 3300 m, 28.8.1954, Polunin sub Davis 24762b (E). — B9 VAN: Distr. 
Baskale, Ispiriz Dag, 2800 m, 31.7.1954, Polunin sub Davis 23640 (E, M); Artos Dagh, N slopes above Gevas, 2900 m, 
2.9.1956, McNeil 775 (E).
IRAN: PROV. SEMNAN: Damghan-Semnan, im Kuh-i Schahmirzad, östlich Gaduk, 2500 m, 1.8.1948, Behboudi & Aellen 1038 
(W); Shahrud, Kuh-e Abr, Kuh-e Ghatri, 2600 m, 23.6.1974, Wendelbo & Foroughi 12945 (W); Shahrud, in saxosis calc. 
jugi Abr a Shahrud boreo-orientem versus, 1950-2500 m, 22.5.1977, Rechinger 56346 (B, M). — PROV. GORGAN: In 
declivibus borealibus montium Shahvar prope Hadjilang, Gardane Dasht, 26.-27.7.1948, K.H. & F. Rechinger 6221 (E, G, 
W); ibid., ad Astamaidan, 3200 m, 26.-27.7.1948, K.H. & F. Rechinger 6043 (B, G, M, W); Cehel-Dokhtar, 6.1948, Sharif 
152 (W). — PROV. MAZANDARAN: Central Alborz, Ilekah road 4 km above Pol-e Zanguleh, 2420 m, 7.10.1967, Baker 74 
(W); m. Elburs occid. supra pagum Paratschan, ad. rud. m. Takht-i-Soleiman (ad Kaki), 2800-3000 m, 28.6.1902, J. & A. 
Bornmüller 6905 (B); Elbursgebirge, Nordhänge, Sardabrud-Tal gegen den Takht-e Suleiman, 1600-4400 m, 7.1977, Cramer 
VO-77/419 (MSB); Kalardasht, Takhte Soleyman montis, 2550-3950 m, 13.9.1970, Termeh 14649 (W); Elika, Kamarbon to 
Varvasht, 2500-3200 m, 15.7.1980, Termeh & al. 39833 (W); Kalej. Meli-Poshteh, Kouhaye Sardabe, 2250-2750 m, 
8.7.1982, Termeh & al. 41132 (W); distr. Nur, inter Kamarband et jugum Naftab, 2600-3200 m, 8.8.1948, K. H. & F. 
Rechinger 6466 (G, W). — PROV. TEHRAN: Chalus road, N facing slopes of Kandavan pass, 2400 m, 21.6.1974, Alava 13994 
(TUR); on the pass between Tehran and Chalus, 2700 m, 3.7.1969, Andersen & Petersen 190 (C, K, W); Kandovan, Chalus 
valley, 2970 m, 21.7.1971, Ahmadi 2116 (LE); 31 km E Firuzkuh, Kadok, 2100 m, 11.6.1972, Babakhanlu & al. (W); Karaj-
Chalus, Kandavan, 2220 m, 28.6.1973, Babakhanlu & Amin 15414 (W); ibid., 2550 m, Babakhanlu & Amin 15023 (W); 
Karaj-Chalus, exclosure Pol-e Zanguleh, 2400 m, 24.7.1973, Babakhanlu & Amin 15242 (W); Karaj-Chalus, Gajereh, 2500 
m, 7.9.1973, Babakhanlu & al. 15542 (W); M. Elburs occid., in reg. alpina jugi Kandevan, 3000 m, 23.6.1902, J. & A. 
Bornmüller 6908 (B, K, LE, W); M. Elburs occid., in alpina Gardene Barry ditionis Assadbar, 3000 m, 26.6.1902, J. & A. 
Bornmüller 6909 (B); M. Elburs occid., in reg. alpina inter Dschoistan et Hasartschal, 3000 m, 28.6.1902, J. & A. Bornmüller 
6905 (B); in regione alpina Totschal, 3500 m, 7.7.1902, J. & A. Bornmüller 6910 (B); Elburz, Gochisar [Gachsar], 50' Ne 
Karaj, 8000', 21.6.1962, Furse 2744 (E, K, W); Elburs, Kandevan, 1935, Gauba 443 (B); Nordhang des Kandevan, 2400 m, 
9.6.1937, Gauba 1479 (B); Elbursgebirge, beim Kandewan pass, 3140 m, 4.7.1936, Gilli 1750 (W); prope Gattade in valle 
Talagon, montium Elburs, 16.7.1843, Kotschy 550 (W); ad pagum Schahristanek prope Teheran in montibus Elburs, 
16.7.1843, Kotschy 550 (W); Elbursgebirge, Südhänge, Varangrud-Tal bei Gachsar, 2500-3700 m, 7.1977, Cramer VO-
77/743 (MSB); E du col de Kandevan, 2900-3050 m, 26.7.1960, Pabot 4486 (G, W); N col Kandavan, 2.6 km S Siah-Bisheh, 
2180 m, 27.7.1960, Pabot 4657 (G); Montes Elburs centr., in jugo Kandawan, in declivibus, c. 2700 m, 9.6.1937, Rechinger 
969 (K, W); in decliv. borealibus jugi Kandavan, 2400 m, 21.6.1974, Rechinger 48284 (W); in decliv. australibus jugi 
Kandavan, 2400 m, 25.6.1975, Renz in hb. Rechinger 53793 (M, W); Shahrestanak, Tuchal, 3100 m, 18.7.1972, Riazi 6143 
(E, LE); Kuhé Totchal, Tchaman-Makhmal to Galeh-Killeh, 2500-3500 m, 2.7.1973, Termeh & Satei 15563 (W); Kandavan, 
Kuh-e Tunnel, 2600-3000 m, 29.7.1983, Termeh & Zargani 41142 (W); Chalus Tal, Siah Bisheh, Sabeti 12414 (W); Montes 
Elborz, in pylis Kandavan in pratis ad meridem vergentibus, 2700-3800 m, 8.7.1977, Sojak 7790 (PR); Kandevan pass, N 
side, 2700 m, 23.6.1959, Wendelbo 2104 (BG, W). — PROV. GILAN: Amarlou, Pak-Deh, 1800 m, 29.6.1967, Darya-Del 
12307 (W); Amarlou, Damash, Kuhé Zard-Tchin, 1750-2250 m, 4.7.1972, Termeh & Daryadel 15225 (W); Amarlou, Kouhé 
Dolfak, 2600 m, 7.7.1972, Termeh 15237 (W). — PROV. GILAN/E AZARBAIJAN: Astara to Ardebil, pass Heyran, 1500 m, 
18.5.1971, Iranshahr 41034 (W); in latere occid. montium Talysch, in valle Ambrani, Ardebil versus, 24.6.1880, Radde 11 
(LE). — PROV. E AZARBAIJAN: Gounas mt., nr. Ardebil, 7800', 17.7.1959, Brown 2116B (W); Sahand Gebirge, 
19.6.1847[1850], Buhse 506 (G-BOIS); Ardebil, 42 km to Nohour, Lisar protected area, 2540 m, 23.7.1974, Foroughi & 
Assadi 13804 (W); 5 km E Kandujan (= 33 km E Kkhosrovshah), on Sahand mt-mass, 2900 m, 19.7.1964, Grant 16282 (W); 
Isperechan in m. Sehend elatioribus, 4.8.1844, Knapp (WU: possibly an isotype of var. glabrescens); Qotur Su, hot spring 
village at north side of Kuhaye Sabalan SE of Khiyav (Meshkin Shahr), 2300-2800 m, 13.7.1971, Lamond 4668 (E); Montes 
Sabalan, in decliv. borealibus saxosis supra pag. Qotur-Su, 2300-2800 m, 13.7.1971, Lamond & Termeh 44079 (W); Sardabé 
to Kuh Sabalan, 24.7.1968, Termeh 13263 (W); Ardabil, Ghotour, Kouh-e Sabalan, 2200-2800 m, 13.7.1971, Termeh 41017 
(W); Bostanabad, Atmishalti, Sahand, Kouh-e Darvish, 2700-3100 m, 4.8.1984, Termeh & Moussavi 41384 (W); in 
declivibus occident. jugi inter Heyran et Ardabil, 1400 m, 18.5.1971, Rechinger 39934 (M, W); in montibus inter Asalem 
(Navrud) et Khalkhal (Herowabad), supra limitem fagetorum, 1900-2300 m, 16.7.1971, Rechinger 43467 (W); Kiyamaki 
dagh ad boreo-orientem a pago Miab, 2500-2600 m, 17.6.1977, Rechinger 56858 (M, W); Tabriz, Kuhi Saudan, near Meyar, 
2600 m, 29.1.1968, Rix & al. 927 (E); versant N de la Kuh-i Savelan, au dessus de Mishgin Sar, 1600-3550 m, 22.-26.8.1956, 
Schmid 6534 (W); Maragheh, Kuh Sahand, 21.6.1965, Sharif 6326 (W); Gardaneh Almas, highest pass on the road from 
Assalem to Khalkhal, 2350-2400 m, 16.7.1975, Wendelbo & Assadi 18486 (LE, W). — PROV. W AZARBAIJAN: Kuh Kani 
Ziarat N Habashi Bala prope Qotur, 18.7.1974, Rechinger & Renz 49642 (M, MSB, W).
NOT TRACED: Aderbaijan, Aucher-Aloy 4395 (G, G-BOIS, LE, P, W); Subalp. Armenia, Aucher 1254 (G, G-BOIS, P); Persia, 
Elburs, Gauba 962 (B); Persia, in montis Karnaru lapidosis, 6.10.1884, Knapp (B: not the isotype of A. caspicus var. 
knappianus Širj.); Dagestan, 8.1876, Becker (G-BOIS, P); Persia borealis, 1836, Ledebauer (P); Iran, prov. Yehlah, Tspili, 
1936, Lindsay 1295 (BM); Georgia Caucasus, Don de Pierre Marten (BR); in alpe Tufan dagh, Caucasus orientalis, 
31.7.1830, Meyer 1265 (G-Bois, LE); Caucasus, Meyer (P, WU); Diabar, 5.1870, Radde 210/247 (LE); Armenia rossica, 
Seidlitz (P); Armenia, Steven (P); Cauc. orientalis, 1821, Steven (G-DC); Caucasus, 1833, Steven (BR); Asia minor, 1858, 
Tchihatchef 892 (G-BOIS, P).


4. Astragalus bactrianus Bunge

UZBEKISTAN: M. Agalic, prope Samarcande, 29.7.1897, Jaccard (ZT).
TADZHIKISTAN: Sarafshan [Zeravshan], Ouranmitane, 15.6.1881, Capus 351 (LE: see the note under A. asaphes); in valle 
Sarawschan, ad ripas m. prope pag. Gusar [Gisar], 1240 m, 18.7.1913, Fedtschenko (in Bornmüller 140); Alpes Sarawschan, 
in decliviis montium aridis prope p. Kischtut, c. 1520 m, 29.7.1913, Fedtschenko (in Bornmüller 149: B, LE); Seravchanica, 
Kum, 3.6.1893, Komarov (LE); ibid., 20.7.1913, Kükenthal (in Bornmüller 96, LE); Sarawschan, inter Gusar et Kschtut, 
3000-4000', 18.-20.6.1882, Regel (LE).


5. Astragalus beckerianus Trautv.

ARMENIA: In latere meridionali montis Aragac, secus viam ad lacum Kari ducentem, 2900 m, 13.7.1975, Gabrielian 12828 
(G).


6. Astragalus brachycalyx Fisch.
6a. Astragalus brachycalyx subsp. brachycalyx 

TURKEY: B8 BITLIS: In m. Meleto Dagh (Meretung Dag) districtus Bitlis, 1800-3100 m, 10.-11.8.1910, Handel-Mazzetti 2776 
(W, WU). — B8 SIIRT: Sasun daglari, Halkis Dag, N & NW slopes above Sason, 1600 m, 22.7.1967, Watson & al. 3459 (E). 
— C9 HAKKARI: Cilo daglari, Zentralkurdischer Taurus, 15.-18.8.1957, Deutschmann BO/19 (WU); Uldere, Süvari Halil, 
2500-2600 m, 26.7.1983, Ekim 8064 (GAZI); Cilo Dag, above Diz deresi, 2000', 7.8.1954, Polunin 23955 (E, K); Cilo Dag, 
below Cilo yayöa, 9000 m, 7.8.1954, Polunin 23974 (E).
IRAQ: MAM: Dohoki, nr. Sersang, 5500', 9.8.1957, Haines 1206 (E); ibid., 4000', 14.7.1955, Haines 573 (E). — MRO: Khani 
Pirbasa, 7900', 9.8.1956, Haley 78 (BM); Warshanka-Margar range, 2200-2500 m, 25.8.1957, Rawi & Serhang 24334 (K); 
Qarnaqo valley, N Pushtashan, 1000 m, Rawi & Serhang 26603 (K); distr. Erbil, m. Qandil in declivibus orient. supra 
Pushtashan, c. 1800-2200 m, 28.7.-1.8.1957, Rechinger 11761 (G, W); distr. Erbil, m. Helgurd in valle supra pagum 
Nowanda, c. 2600-3000 m, 10.-14.8.1957, Rechinger 11464 (G, W). — MSU: Suleimanya, Hero, 1800 m, 17.7.1947, Rawi 
9384 (US).
IRAN: PROV. W AZARBAIJAN: Oroumieh, Razhan, Khalil Kuh, 1600-1800 m, 9.7.1994, Chehregani & Zarre 17876 (MSB, 
TARI, TUH); 34.5 km S Mahabad, 1720 m, 16.10.1960, Pabot 5492 (G); ibid., 1840-1930 m, Pabot 5508 (E, G, MSB, W); 
Chalil Kuh, in faucibus N Razhan, 1800-2000 m, 2.7.1974, Rechinger 48753 (W); Chalil Kuh, in faucibus NW Selvana, 
1750-2000 m, 4.7.1974, Rechinger 48924 (W). — PROV. KERMANSHAH: 6 km after Paveh on the road to Ravansar, 1600-
1700 m, 7.7.1994, Chehregani & Zarre 17832 (M, TARI, TUH). — PROV.KORDESTAN: Above Sardasht, 6000', 29.7.1962, 
Furse 3493 (K, W). — PROV. BAKHTIARI: 34 km to Chehelgerd from Farsan, 2 km after Heydar Abad, 17.7.1994, Zarre 
17777 (MSB, TARI, TUH). — PROV. ESFAHAN: Balehsun Kuh, between Damaneh and Khunsar, 2740 m, 14.7.1966, 
Archibald 2715 (E, K); Golpayegan-Khonsar, Golestan Kuh, 2800-3250 m, 1.7.1969, Iranshahr 14698 (W); Nadjun, 1900-
2200 m, 14.5.1937, Köie 592 (B, C); 30 km to Daran from Esfahan, Dallan Kuh, Cheshmeh Gorgi, 2280-2500 m, 2.8.1973, 
Moussavi & Satei 15537 (W); Khonsar, Ghaleh Baba-Mohammad, Golestan Kuh, 2480-2850 m 9.7.1984, Moussavi & 
Tehrani 41586 (W); in montibus pr. Damaneh 35 km SE Daran, 115 km NW Esfahan, 10.6.1974, Rechinger 47663 (W); 
Khonsar, 1640 m, 31.7.1970, Riazi 2253 (E, LE); Sultanabad ad Chonsar, 12.8.1892, Strauss (B, JE). — PROV. LORESTAN: 
Azna, Bavaki, 2200-2800 m, 14.7.1969, Iranshahr 14700 (W); Dorud, 5000', 2.6.1941, Koelz 18512 and 28.6.1941, Koelz 
18387 (W); in m. Schütrün Kuh [Oshtoran Kuh], 7.1890, Strauss (B, JE, K, L, WU); ibid., 28.7.1902, Strauss (G); ibid., 
8.1903, Strauss (B, JE, W).


6b. Astragalus brachycalyx subsp. eriostylus (Boiss. & Hausskn.) Zarre

IRAN: PROV. BAKHTIARI: Kuh Cherri, 5.6.1899, Escalera 67286 (MA); ibid., 7.1899, Escalera 67287 (MA); Ardal, Zard Kuh, 
2700-3200 m, 14.-15.6.1973, Iranshahr & Moussavi 15598 (W); Ardal, Chery to Dashtak, 2100 m, 16.6.1973, Iranshahr & 
Moussavi 15572 (W); Barrage de Kuh-Rang, 2500 m, 6.7.1959, Pabot 2157 (G); ibid., 12477 (W); 80 km S Gandoman, 2300 
m, 9.7.1959, Pabot 2295 (G); Borujen to Lordegan, 42 km from Borujen, 2600 m, 14.9.1993, Zarre 72062 (MSB, TARI, 
TUH). — PROV. LORESTAN: Khorram-Abad, Taf area, 10.5.1970, Baharvand 140163 (Shiraz); Kuh Pirhashtad, 12.8.1950, 
Hakim 5245 (W); Sarab Tak, 2100-2500 m, 8.1982, Mossadegh 3 (E); Karon bei Bisheh (route Dizful-Sultanabad), 2200 m, 
5.6.1937, Köie 128 (B). — PROV. FARS: Shiraz, Tolé Khosrow, Kakan, Kuh-e Kalivar, 23.7.1949, Behboudi 1076 (W); near 
Dasht-e Arjan, about 70 km W Shiraz, on road to Kazeroun, 2000 m, 1.6.1972, Bokhari 652 (Shiraz); Ghalat, 35 km NW 
Shiraz, 2300 m, 8.6.1965, Grant 17708 (Shiraz, W); 76 km NW Ardakan, 2300 m, 14.7.1959, Pabot XLIVC (G).


7. Astragalus breviflorus DC.
7a. Astragalus breviflorus subsp. breviflorus 

TURKEY: B8 ERZURUM: Ad Erzerum, 1834, Aucher-Aloy 2506 (W); Palandöken Dag, 20-23 km from Çat to Erzurum, 2300-
2350 m, 27.7.1966, Davis 47357 (E); Erzurum-Çat, bei Tasligüneg, 2200 m, 18.8.1987, Engel 142 (MSB); in montibus Tech-
Dagh supra Erzeroum, 7000-8000', 7.1853, Huet Du Pavillon (E, G, G-BOIS, P); Erzurum - Çat, 25 km N Çat, 2400 m, 
13.7.1988, Nydegger 43722 (MSB); Erzurum - Bingöl, 2250 m, 29.6.1992, Nydegger 46518 (MSB). — B9 BITLIS: 
Adilcevaz, Harmanteppe village, Süphan Dagi, c. 3000 m, 7.8.1989, Aytaç & Metzger 2982 (GAZI).
NOT TRACED: Armenia, Aucher-Aloy 1256 (G, G-BOIS).


7b. Astragalus breviflorus subsp. elongatus (D.F. Chamb. & V.A. Matthews) Zarre

TURKEY: B9 BITLIS: Nemrut Dag, 7600', 3.7.1954, Polunin 23572 (E); ibid., S exposition, 2250 m, 9.8.1987, Engel 098 
(MSB).


8. Astragalus carduchorum Boiss. & Hauskn.

IRAQ: MRO: Riwandous ad fines Pers., supra monte Sakri-Sakran, 1800 m, 23.6.1893, Bornmüller 11197 (B); Erbil liwa, 
valley E Arl Gird Dagh, 2100 m, 4.8.1947, Gillet 9488 (K); Rayat, 2300 m, 25.8.1953, Guest 13115 (K); Pir Omar Gudrun, 
ad nives 12000', 6.1867, Haussknecht (G); Erbil liwa, Mergadreija near Haji Omran, 1700-1800 m, 21.6.1947, Rawi 9168 
(K); Helgurd, 1800 m, 1.7.1954, Rawi 13794 (K); Bask, Haworan, c. 1500-1800 m, 29.7.1957, Rawi & Serhang 23959 A (K); 
Warshanka-Magar range, c. 2200-2500 m, 25.8.1957, Rawi & Serhang 24328 (K); lower slope of Qandil range, c. 1450 m, 
18.8.1958, Rawi & Serhang 26706 (K); distr. Erbil, Montes Qandil ad confines Persiae, in decl. orientalibus supra 
Pushtashan, 1800-2200 m, 28.7.-1.8.1957, Rechinger 11077 (B, E, G: see the note under A. chionobiiformis).
IRAN: PROV. KORDESTAN: C. 90 km (E side of the pass), from Sanandadj to Marivan (Dezh Shahpur), c. 1650-1800 m, 
6.7.1971, Lamond 4536 (E); Sanandaj to Marivan, 90 km to Sanandaj, 1650 m, 5.7.1971, Termeh 14770 (W); in querceris 
(Qu. brantii) 90-110 km W Sanandaj versus Dezh Shahpur, 1650-1800 m, 6.7.1971, Rechinger 42948 (B, G, K, W); Montes 
Chahel Chashmeh, 44 km NE Marivan, 2000 m, 7.7.1971, Rechinger 43047 (G, W).


9. Astragalus caspicus M. Bieb.
9a. Astragalus caspicus subsp. caspicus 

AZERBAIJAN: Gub. Kuba, in siccis supra p. Gümur, 4500', 11.7.1899, Alexeenko 1889a (LE); ibid., 5000', Alexeenko 1773 
(LE); Gub Kuba, ad fl. Ata-czai infra p. Alty-agacz, 3500', 24.7.1900, Alexeenko 5904 (LE); Gub. Kuba, distr. Schemacha, ad 
fl. Czikal-czai, inter Alty-agacz et Schichlar, 2600', 26.7.1900, Alexeenko 6649 (LE); ibid., 3600', 24.7.1900, Alexeenko 5947 
(LE); ibid., pr. p. Chilmil, 3000', 27.7.1900, Alexeenko 6691 (LE); ibid., in pascuis montis Dibrar, 6200', 25.7.1900, 
Alexeenko (LE); Baku, Becker (P, WU); Caucasus, Maraza, 11.6.1952, Al. Federov & al. (LE); Swant, prope pag. Schon 
czola, 7.1952, An. Federov & Rzazade (LE); prope pag. Alty-agacz, 9.8.1952, An.Federov & Smoljiaminov (LE); prope 
Baku, 1863, Geubel (G-BOIS); in collibus aridis arenosis montium ditionis Swant Georg. Cauc. 4000'-6000', 6.1836, 
Hohenacker (E, G-BOIS, L, LE, M, P, W, WU); Amir-Alag, na Zukhikh sklona, 1400 m, 10.7.1960, Kharedi & Ivan (E); 
Dyman near Baku, 20.5.1898, Lewandowsky (LE); in collibus lapidosis pr. pag. Swant, 20.6.1830, Meyer 1366 LE); Baku, 
1842, Meyer 1266 (G-BOIS, P); Talish, 8.1844, Meyer [254] (LE, W).
TURKEY: B9 VAN: Van, Kampüs, 1700 m, 29.9.1992, Özçelik 4719 (GAZI).
IRAQ: MRO: Riwandous ad fines Pers., in monte Sakri Sakran, 1900 m, 23.6.1893, Bornmüller 1198 (B, JE). — MAM: Distr. 
Mosul, ad confines Turciae prov. Hakkari, inter Dohuk et Amadiya, in quercetis infra Sirsank, 10.-12.7.1957, Rechinger 
11612 (G, M, W); Sersang, open stony slope in valley bottom, 4000', 5.8.1957, Wheeler Haines (E). — MSU: Pir Omar 
Gudrun, 4000-6000', 6.-7.1867/68, Haussknecht 346 (BM, G-BOIS, JE, LE, M, MSB, P, W); Suleimanya, Hero, 1800 m, 
17.7.1947, Rawi 9387 (US).
IRAN: PROV. SEMNAN: 20 km SW of Fulad Mahalleh, 1900 m, 15.6.1973, Andersen & Jensen 7007 (C, E, L); 35 km 
Shahmirzad to Fulad Mahalleh, 2350-2500 m, 9.7.1974, Renz & Iranshahr 16732 (W). — PROV. TEHRAN: 7 km E Firuzkuh, 
Shahdasht, 1900 m, 10.7.1972, Babakhanlou & al. 16052 (W); Rostamabad, road of Firuzkuh, 2550 m, 27.6.1972, Dini & 
Arazm 15599 (W); Ozineh, road of Firuzkuh, 2310 m, 5.7.1972, Dini & Arazm 14734 (W); road of Firuzkuh, 2400 m, 
8.7.1972, Dini & Arazm 15835 (W); Firuzkuh, 2000 m, 25.7.1972, Dini & Arazm 15101 (W); 20-36 km on the road from 
Damavand to Firouzkuh, 29.8.1993, 2100 m, Maassoumi & Zarre 71928 (MSB, TARI); 15.5 km E Teheran, 1460 m, 
9.7.1960, Pabot 4010 (G); 115-120 km E Teheran, plateau SW de Firuzkuh, 1950-2150 m, 12.7.1960, Pabot 4350 (G); 75 
km E Teheran, Station de Homand, 1900 m, 13.6.1965, Pabot 7447 (G, W: sub 12478); 22 km from Firuzkuh towards 
Rudhen, near Chehel Cheshmeh, 2300 m, 12.7.1974, Renz & Iranshahr 16727 (W). — PROV. GILAN: W du barrage du Sefid-
Rud, 350 m, 10.6.1960, Pabot 3652 (G); Manjil, above Tunnel, 370-400 m, 22.8.1993, Zarre 72050 (MSB, TARI, TUH). — 
PROV. ZANJAN: Zanjan-Hamadan road, hillsides and meadows along the irrigation canal about 7 km from Soltanieh on the 
road to Qeydar, 1850 m, 2.7.1974, Alava 14181 (TUR); Zanjan-Hamadan road, roadside fields and roadbanks at the village 
Arghin, about 30 km from Soltanieh on the road of Qeydar, 1700-1800 m, 2.7.1974, Alava 14201 (TUR); Zanjan-Hamadan 
road, rocky hillsides between the village of Arghin and Dashti at the foot of Qeydar mountain, about 35 km from Soltanieh, 
1980-2200 m, 2.7.1974, Alava 14230 (TUR); Kaswin, 6.1859, Bunge & Bienert (P); inter Chorumderreh et Sfiud, inter 
Teheran et Tabris, 6.1859, Bunge & Bienert (P); 80 km S Qazvin on the road of Zanjan, 1640 m, 4.7.1972, Foroughian 
16034 (W); 84 km SW Qazvin, 1680 m, 9.7.1972, Foroughian & Hariri 15964 (W); Sultanabad, near Kasvin, 5000', 5.1935, 
Lindsay 254 (BM); 50 km SE Zanjan, S Soltanyeh, 1800 m, 24.6.1960, Pabot 3905 (G); ad versuras 80 km NW Takestan 
versus Zanjan, 1500 m, 29.6.1971, Rechinger 42341 (M, W); Zanjan, in collibus argilloso-conglomeraticis, 1600 m, 
13.6.1977, Rechinger 56515 (W); Ghazvin to Zanjan, 29.6.1971, Termeh 40790 (W); Soltanieh, 7 km to Gheydar, 1850 m, 
2.7.1974, Termeh & Moussavi 16467 and 40805 (W). — PROV. E AZARBAIJAN: Pr. Nehmatabad, inter Teheran et Tabris, 
6.1859, Bunge & Bienert (P); 10 km E Sarab, 5000', 14.6.1929, Cowan & Darlington 1598 (K); near Tabriz, 1927, Gilliat-
Smith 1905b (K!); 5 km E Kandujan (= 33 km E Khosrowshah), on Sahand mt-mass, 2900 m, 19.7.1964, Grant 16,281 (W); 
prope Tabris, 1350-1500 m, 25.6.1924, Grossheim (B); 10 km N Marand, 1600 m, 6.6.1971, Iranshahr 14753 (W); red hills 
N Tabriz, c. 1700 m, 20.7.1971, Lamond 5182 (E); 6.3 km SE Tabriz, 1500 m, 8.10.1960, Pabot 5154 (G); 5 km S Marand, 
1580 m, 11.10.1960, Pabot 5323 (G); supra Marand, 1400 m, 6.6.1971, Rechinger 41159 (W); in jugo inter Marand et 
Sufian, 1600-1750 m, 6.6.1971, Rechinger 41182 (M, W); in decliv. siccis et ad versuras prope Amirabad inter Mianeh et 
Khalkhal (Herowabad), 1800-2000 m, 14.7.1971, Rechinger 43307 (W); in collibus rubris ‘Einal Zeinal’ inter Tabriz et 
fluvium Talkheh Rud (Atschi Tschai), 1700 m, 20.7.1971, Rechinger 43557 (W); in decliv. siccis inter Sufian et Shabestar, 
1400 m, 22.7.1971, Rechinger 43598 (W); ad versuras lapidosis infra Kandavan versus Usku, 2000 m, 20.6.1977, Rechinger 
57136 W); environs de Ardebil, 1000-1600 m, 20.-21.8.1956, Schmid 6498 (W). — PROV. W AZARBAIJAN: 45 km to 
Uroumieh from Salmas, Ghoushchi pass, 8.1994, Chehregani 17945 (MSB); Oroumieh, Razhan, Khalil-Kuh, 1600-1800 m, 
9.7.1994, Chehregani & Zarre 17847 (M, TARI, TUH); Rezaieh, 32 km de Band vers Moana, 1550-1750 m, 2.6.1978, Matin 
& Daneshpajuh 38406 (W); in collibus argilloso 30 km S Khovy versus Shahpur, 1500 m, 12.6.1971, Rechinger 41799 (W); 
a Rezaiyeh 26 km versus Sero, 1600-1700 m, 14.6.1971, Rechinger 42059 (M, W); in colle argilloso SW Rezaieh, 1500 m, 
12.7.-13.7.1974, Rechinger 49310 (W); inter Ashgharabad et Kolavana NNW Khovy, 2100 m, 16.7.1974, Rechinger 49434 
(M, W); in jugo Qushchi inter Shahpur et Rezayeh, 1900 m, 21.7.1974, Rechinger 49809, 49866 and 49867 (W) in montibus 
ad limites Turcia, distr. Khoi, 17.7.1828, Szovits 542 (G-BOIS, LE, P, W). — PROV. KORDESTAN: Boukan, 1.8.1964, 
Haghighi 6259 (W); Saqqez to Tabriz, c. 11 km N of Saqqez, 1500 m, 8.7.1971, Lamond 4618 (E, M); 26 km E Sanandaj, c. 
2200 m, 28.6.1965, Ledingham & al. 4240 (W); in saxosis (Tonschiefer) 6 km SE Bijar, ad viam versus Hamadan ducentem, 
2000 m, 30.6.1971, Rechinger 42747 (W); Kuh-e Hamzeh Arab, Bijar to Hamadan, c. 2000 m, 1.7.1971, Rechinger 40395 
(E, W); in collibus siccis et ad versuras 11 km N Saqqez, 1350 m, 8.7.1971, Rechinger 43103 (W); ad versuras 33 km NW 
Divandarreh versus Saqqez, 2100 m, 29.6.1974, Rechinger 48590 (W). — PROV. HAMADAN: Yengedjeh, 20.7.1960, 3000', 
Brown 2954A (W); Razan to Avaj, 10 km to Avaj, on the neck, 2200-2300 m, 10.7.1994, Chehregani & Zarre (M, TARI, 
TUH); 53 km W Hamadan, 1850 m, 19.10.1960, Pabot 5578 (G); col entre Hamadan et Qazvin, 2200 m, 20.10.1960, Pabot 
5588 (G); Montes Karaghan, in jugo Soltan Bolagh inter Avej et Razan, 2200-2400 m, 27.6.1974, Rechinger 48458 (M, W); 
ad versuras inter Qorreh et Dehgolan, 120 km ab Hamadan versus Sanandaj, 1950 m, 28.6.1974, Rechinger 48469 (W); 
region d’Aq Bulagh, 90 km N Hamadan, 2000 m, Rioux & Golvan AB288 and AB289 (G, W). — PROV. KERMANSHAH: 7 km 
from Paveh to Ravansar, 1700 m, 7.7.1994, Chehregani & Zarre 17839 (MSB, TARI, TUH). — PROV. ARAK: Between Arak 
and Khomain, Varche village, 2050 m, 15.6.1984, Mozaffarian & Maassoumi 48022 (MSB, TARI); 6 km W d’Arak, 1920 m, 
7.6.1959, Pabot 1174 (G); 15 km W d’Arak, 2050 m, 7.6.1959, Pabot 1202 (G). — PROV. LORESTAN: 55 km from Arak to 
Malayer, 2100 m, 15.6.1984, Mozaffarian & Maassoumi 48081 (MSB, TARI).
NOT TRACED: Persia in mont Ghilan, Aucher 4392 (G-BOIS, LE, P, W); Kuschkek, 19.6.1882, Pichler 176 (WU); Persia 
borealis, Ledebour (P); Transcaucasia, Szovits (JE).


9b. Astragalus caspicus subsp. pseudocaspius (Fisch.) Zarre

TURKEY: A4/5 KASTAMONU: Tasköprü to Gökçeagaç, 500 m, 24.7.1962, Davis & al. 38061 (E). — A5 AMASIA: In lapidosis 
rupestribus a regione inferiore Calida et alpinam usque copiose, 5.5.1889, Bornmüller 96 (B, BR, G, LE, PRC, WU); in 
subalpinis mt. Ak-dagh, 1000 m, 4.9.1890, Bornmüller 2128 (B, G, JE, LE); c. 2 km S Baglarüstü village, 880 m, 19.6.1994, 
Cansaran 479 (GAZI); ± 13 km SW of Havza along the road Samsun - Çorum, ± 1000 m, 13.6.1959, Hennipman & al. 1958 
(L). — B4 ANKARA: Ankara-Beypazari, c. 1000 m, 3.7.1971, Akman 942 (E); Mihaliçik, 700 m, 8.7.1962, Davis & Coode 
37215 (E).


10. Astragalus caucasicus Pall.

RUSSIAN CAUCASUS: DAGESTAN: Sufun bei Kizil-jurt, am Sulak, Lagowski (LE); bei Derbent, Lagowski 9559 (LE); in locis 
lapidosis ad torrentem Terek, Meyer 1261 (LE).
AZERBAIJAN: Gub. Baku, distr. Kuba, in siccis supra p. Gümur, 5000', 11.7.1899, Alexeenko 1773 and 4500', Alexeenko 
1889a (LE); Gub. Baku, distr. Kuba, ad fl. Ata-czai, infra pag. Alty-agacz, 3500', 24.7.1900, Alexeenko 5904 (LE); Gub. 
Baku, distr. Kuba, ad fl. Czikal-czai inter p. Alty-agacz et Schichlar, 2600', 26.7.1900, Alexeenko 6649 (LE); pr. Baku, 
Eischwald (LE, P); in district. Schamchoz, in angust. fl. Andasa, 19.8.1952, An. Federov & Rzazade (LE); Schamkhor, 1200 
m, 5.7.1956, Gadjiev (LE); in sterilibus collibus deserti ditionis Elisabethpol, 7.-8.1834, Hohenacker (BR, E, G, G-BOISS, 
GOET, L, LE, M, P, W); in aridis saxosis prope Helendorf et in ditione Swant Georg., 6.-7.1838, Hohenacker (G-BOIS, LE, 
M: mixed with A. microcephalus); prov. Gandzha, distr. Kazach, inter St. viae ferr. Salogly et montem Kuhkuna, 14.7.1928, 
Kalkovsky (LE).
GRUZIA: Tiflis, Bongard 142 (E); pr. Tiflis, 6.1881, A.H. & V.F. Brotherus 274 (BR, LE, G-BOIS, P); Tbilisi, hillsides near 
the Dabahane gorge, opposite the Tbilisi Botanical Institute, 550 m, 29.6.1959, Davis 33806 (E, LE); Tiflis, fl. Dabachaika, 
9.9.1918, Grossheim (LE); prov. Tiflis, distr. Signach, in jugo Katari prope p. Uljanovka, in lapidosis, 14.7.1926, Grossheim 
389 (K, LE); in saxosis aridis Gruzia, 8.1838, Hohenacker (GOET, M: mixed with A. microcephalus); Tiflis, Navtlugh, 
9.6.1912, Holmberg 1412 (W); Tiflisia, 29.7.1903, Poon (LE); auf Hügeln bei Tiflis, 1873, Rehman 225 (G-BOIS); Tiflis, 
gori Kruanisi, 29.6.1903, Selezhinskago (E); Okrestnosti Tiflisa, Arsenalnaja, 7.9.1919, Shishkin (LE); reg. pr. Tiflis, 
18.7.1895, Sintenis (L); Tiflis, in collibus aridis ad Merid, montis Davidii, 9.7.1890, Sommier & Levier 347 (B, LE, WU); 
Iberia et Tiflis, Steven (P); peripheria sepentrionales urbis Tbilisi, loco Saburtalo dicto, 500-600 m, 19.7.1988, Vašák (W); 
distr. Khashuri, in vicinitate pagi Igoeti, c. 600 m, 21.7.1988, Vašák (W); Iberia et prov. Tchalziz, 1843, Wittmann (JE).
TURKEY: A9 KARS: Bei Kars, Lagowski 9558 (LE).
NOT TRACED: Caucasus, Bentham 142 (E); Iberia, prope Marienfeld in collibus siccis, Hohenacker (LE); Caucasus, 1836, 
Ledebour (P); Iberia, Meyer (PR!); Somchetia, Meyer (GOET).


11. Astragalus chartostegius Boiss. & Hauskn.

IRAN: PROV. BAKHTIARI: Gandomkar, Kuh-e Garreh, 2300-2700 m, 9.6.1973, Iranshahr & Moussavi 15553, 15554, 15556 
(W). — Prov. FARS: Declivia austro-occidentalia montium Kuh-e Dena, supra vicum Sisakht, 3400-4000 m, 14.6.1973, Sojak 
6323 (PR); Sisakht, Dena, Kuh-e Gol, 3200 m, 18.6.1969, Termeh & Izadyar 14691 (W).


12. Astragalus chionobiiformis C.C. Towns.

IRAQ: MRO: Distr. Erbil, Montes Qandil ad confines Persiae, in decl. orientalibus supra Pushtashan, 1800-2200 m, 28.7.-
1.8.1957, Rechinger 11077 (K: see the note); ibid., Rechinger 11082 (W, MSB).


13. Astragalus coarctatus Trautv.
Known only from type.


14. Astragalus divaricatus Bunge

AZERBAIJAN: Type material of A. jucundus.
IRAN: PROV E AZARBAIJAN: Sharafkhaneh Tabriz, 1345 m, 4.7.1965, Babakhanlou 19930 (W); Shahbil, Kuh-e Sabalan, 2800 
m, 24.7.1974, Foroughi & Assadi 13856 (W); Mount Sabalan, 11500', 27.7.1961, Harrington 318 (K); Meshkin Shahr, 
Sabalan, 3600 m, 2.8.1970, Izadyar 14648 (W); Monte Sabalan, in declivibus borealibus saxosis, 3500-3700 m, 15.7.1971, 
Lamond & Termeh 44157 (G); ibid., Lamond 4758 (E, M); environs de Ardabil, 1000-1600 m, 20.-21.8.1956, Schmid 6500 
(G); Tabriz - Ardabil, 29 km E Sarab, 2000 m, 6.7.1965, Seraj in hb. PABOT SJXIIIb (G); Ardabil, Ghotur-Sou, Kouhé 
Sabalan, 3400-3600 m, 15.7.1971, Termeh 40819 (W); Sarab, Kouha-ye Narmigh, 2000-2400 m, 9.-10.6.1986, Termeh & 
Daneshpajuh 41365 (W).
NOT TRACED: in Persia prov. Ghilan, Aucher-Aloy 4396 (BM, G, W).


15. Astragalus dolonus (Rassulova & Sharip.) Kamelin.
No material of this species was available to me.


16. Astragalus erythrolepis Boiss.
Known only from the type gathering and the type of the synonym A. leucargyreus.


17. Astragalus gevashensis D. F. Chamb. & V. A. Matthews

TURKEY: B9 BITLIS: Kaz Dag above Kotum, 6500', 28.6.1954, Davis & Polunin 22268 (E, K, M); Hanemir Dag, N Seite von 
Oboskü köyü aus, 2250 m, 10.8.1987, Engel 105 (MSB); Tatvan, Pelli Dag, Syamaç, 2500-2800 m, 19.8.1972, Pesman 3213 
(HUB); in jugo Kuzgunkivan Geçidi inter Gevas et Resadiye, 2200 m, 1.7.1975, Rechinger 53899 (M, W).


18. Astragalus hareftae (Nábelek) Širj.

TURKEY: B9 VAN: Güzel Dere pass, 34 km from Baskale to Hosap, 2800 m, stony slopes in sandy soil, mountain steppe, 
3.7.1966, Davis 45974 (E, M); Van-Hosap, Güzeldere geçiti, 22.8.1993, Y. Altan 5565 (GAZI); dist. Baskale, Ispiriz Dag, 
3400 m, 31.7.1954, Davis & Polunin 23704 (BM).


19. Astragalus hystrix Fisch.
Only known from the type collection.


20. Astragalus icmadophilus Hand.-Mazz.

TURKEY: B7 TUNCELI: Munzur dag above Ovacik, 2700 m, 17.7.1957, Davis & Hedge D.31252 (M); Tunceli, Ovacik, 
Munzur daglari, 2300 m, 28.7.1979, Yildirimli 2237 (HUB). — B9 BITLIS: Hanemir Dag N Seite, von Oboskü Köyü aus, 
2400 m, 10.8.1987, Engel & Eggli 108b (MSB). — B9 VAN: Gürpinar to Baskale, Güzeldere Geçidi , 2850 m, 12.8.1987, 
Engel 117 (MSB). — C10 HAKKARI: Cilo Dag, 10 km W Cilo Tepe, 10300', 9.8.1954, Davis & Polunin 24129 and 24155 
(BM, E); Cilo daglari, Zentralkurdischer Taurus, 24.8.1956, Deutschmann (W); Sat Dag (above Yüksekova), 2800 m, 
26.8.1967, Duncan & Tait 48 (E); Cilo Dag, Avaspi-Gletscher, 2600 m, 17.8.1968, Vogel (M).
IRAQ: MRO: Erbil, Montes Qandil ad confines Persiae, c. 3500 m, 28.7.-1.8.1957, Rechinger 11794 (W).


21. Astragalus iodotropis Boiss. & Hohen.

IRAN: PROV. MAZANDARAN: Damavand, 2750 m, 14.8.1973, Amin 15457 (W); Mt. Elburs, in reg. alpina jugi Demawend, 
3500 m, 16.7.1902, J. & A. Bornmüller 6895 (B); Elbursgebirge, N-Hänge, Sardabrud-Tal gegen den Takht-e Sulaiman, 
1600-4400 m, 7.1980, Cramer VO-77/538 (MSB); Demavend, 9000-12000’, 13.6.1960, Furse & Synge 459 (K); 
Elbursgebirge, Demawend ober Rehne, c. 4000 m, 23.7.1936, Gilli (W); heatwaters of the Chalus river, N slope of Elburz 
mts, 77 km S Chalus, 2800 m, 25.7.1964, Grant 16,464 (W); E slope of Demavend, 7 km SW Reyneh, 4000 m, 29.7.1964, 
Grant 16,539 (W: mixed with A. ochrochlorus); Larijan, Rineh, Gousefand-Sara, (12 km NW Rineh), 3000-3500 m, 
10.6.1980, Moussavi & al. 39857 and 11.9.1980, 39856 (W); Larijan, Rineh, Kuh-e Damavand, 4150 m, 18.6.1982, 
Moussavi & al. 41122 (W); Elika, Kamarbon, Kuh-e Varvasht, 3200-3650 m, 13.-14.7.1980, Termeh & al. 39829 (W); 
Kaledj, Meli-Poshteh, Kouhay-e Sardabe, 2250-2750 m, 8.7.1982, Termeh & al.41131 (W). — Prov. TEHRAN: Dizin, 3400 
m, 31.6.1973, Amin 15010 (W); M. Elburs occid., in reg. alpina inter Dschoistan et Häsartschal (ad Haki), 2800 m, 
28.6.1902, J. & A. Bornmüller 6915 (B); in summis jugis Totschal, 3750 m, 8.7.1902, J. & A. Bornmüller 6916 (B); jugi 
Elbursensis occid. in reg. alpina montis Kandevan, c. 3100 m, 23.6.1902, J. & J. Bornmüller 6917 (B, G, K, W, WU); 
Elburs-Gebirge, Kandevan-Paß und Dalir-Tal, Sommer 1977, Cramer 77/1006 (MSB); Elburz, Gochisar [Gachsar], 50' NE 
Karaj, 8000', 20.6.1962, Furse 2715 (K, W); Elburs, Gauba 961 (B); M. Elburs occid., M. Chalan entre Chalan et Khurram-
Dasht, 3790 m, 7.6.1971, Klein 8265 (MSB); E du col de Kandavan, 2900-3050 m, 26.7.1960, Pabot 4485 (G, MSB); 
Montes Elburz, in saxosis calc. jugi Dizin inter Shemshak et confluentes fluvii Karaj, 3100-3250 m, 25.6.1977, Rechinger 
57256 (G, M, MSB, W); Karadj Tal, Kandavan pass, 6.1959, Sabeti 12423 (W); Tehran to Tchalus, Kandavan, Kuh-e 
Tunnel, 2250-3000 m, 15.7.1972, Termeh 15223 (W); Alborz, Gatchsar to Gajereh, Varang-roud, 2240-2340 m, 8.7.1977, 
Termeh & Matin 36743 (W); Kuh-e Totschal, Tchaman-Makmal to Galeh Kileh, 2500-3500 m, 2.7.1973, Termeh & Satei 
15562 (W); Elburz mts., Kandevan pass, N side, c. 2700 m, 23.6.1959, Wendelbo 2094 (BG).


22. Astragalus karabaghensis Bunge

ARMENIA: Goris, Karashen, 9.7.1952, coll. ignatus (MSB); distr. Karajan, Mt. Meidzhili, 400-500 m, 7.6.1932, Karajagin 
(L); Ekhelnadurekmii raion, Amash-Areni, 11.7.1976, Manakli & al. (LE).
AZERBAIJAN: (NAGORNO-KARABAGH): Karabach, m. Zyarat, 400'-6000', 20.7.1929, Achverdov & Doluchanov in hb. 
GROSSHEIM (B, LE); Vank-Disak, prope pag. Gadrut, in decliv. septentrionalibus, 13.8.1935, Heidemann (LE); distr. 
Karabagh, Medzhili, Ch. d’Alleizette (P); prov. Elisabethpol, Karabagh, inter Kubatly et Dzhebrail, in decliv. arenosis, 
30.5.1911, Vvedensky 386 (B). — NAKHICHEVAN: Norashenskii, Severned, sel. Karabagliar, 20.6.1957, Tzvelev & Czerpanov 
133 (LE).


23. Astragalus kordloricus Zarre

IRAN: PROV. KERMANSHAH: Sarabe Karand, 24.6.1968, Iranshahr 13268 (W); 39 km SW Kermanshah, 1600 m, 14.6.1959, 
Pabot 1860 (G); 44 km W Shahabad, Abiyan-Genk, 1650 m, 14.6.1959, Pabot 1903 (G).


24. Astragalus latus Zarre

IRAQ: MSO, Erbil liwa, Ser Kurawa, 2940 m, 11.8.1947, Gillett 9765 (US).


25. Astragalus leiophyllus Freyn & Bornm.

TURKEY: C10 HAKKARI: Sat Dagi, between Varegöz & Sat Gölü, 26.6.1966, Davis 45542 (E); Kurdistania turcica distr. 
Hakkari, mons Kela Mame supa pagum Hor dit Gulamerik, supra 2500 m, Nabélek 3082 (SAV).
IRAQ: MRO: Erbil liwa, NE slopes of Arl Gird Dagh, 2600 m, 5.8.1947, Gillett 9563 and 2000 m, Gillett 9536 (K); Erbil 
liwa, Ser Kurawa, 2200-2700 m, 11.8.1947, Gillett 9767 (K); NE Qandil, c. 2650-2850 m, 25.8.1957, Rawi & Serhang 24377 
(K: photo MSB); distr. Erbil, mons Helgurd ad confines Persiae, in valle supra pagum Nowanda, 2600-3000 m, 10.-
14.8.1957, Rechinger 11462 (W).
IRAN: PROV. W AZARBAIJAN: Rezaiye [Oroumieh] Qasemlue valley, 1500 m, 1.7.1976, Siami & Zehzad 6982 (K); Rezaiye 
[Oroumieh] Margavar, 2100 m, 3.7.1972, Djavan Moghadam 149 (W).
IRAQ/TURKEY?: Distr. Berwari, Halakur-Dagh, c. 2500 m, 25.7.1910, Nábelek 3072 (SAV).


26. Astragalus macrosemius Boiss. & Hohen.

IRAN: PROV. MAZANDARAN: Elburs occid. in regione alpina montis Takhti Soleiman, ad nives prope Häsartschal, c. 4000-
4100 m, 29.6.1902, Bornmüller 6921 (B, K, W); Elika, Varvasht (montis), 3500-4100 m, 4.8.1972, Termeh 15241 (W); 
Kalardasht, Kuh-e Takht-e Soleyman, 3600-4100 m, 10.8.1972, Termeh 15234 (W); Elika, Kamarbor, Kuh-e Varvasht, 3200-
3650 m, 13.-14.7.1980, Termeh & al. 39832 (W). — Prov. TEHRAN: M. Elburs, in excelsus alpinum Totschal, c. 3700 m, 
8.7.1902, J. & A. Bornmüller 6919 (B) and 6920 (B, G); in sumo jugo alpium Totschal, 3750 m, 4.6.1902, J. & A. 
Bornmüller 6918 (B); prov. sommet Bozorg de Kahar, 3800 m, 18.6.1971, Klein 946 (MSB); Mas Mischenou, 3810 m, 
20.6.1971, Klein 1142a (MSB); Tehran, in declivibus ad pedem montium Elburz supra vicum Darband et declivia montis 
Kuhha-ye Touchal, 2000-3800 m, 29.6.1973, Sojak 7043 (PR); Montes Elborz, in pylis Kandavan in pratis ad meridiem 
vergentibus, 2700-3800 m, 8.7.1977, Sojak 7802 (PR); Iskamar, Kuh Tochal, 2.7.1968, Termeh 13262 (W); Touchal, S side, 
3500 m, 6.9.1974, Wendelbo & Cobham 14495 (W).


27. Astragalus michauxianus Boiss.

TURKEY: C10 HAKKARI: Ciyasipi, Beyaz Dag, NE spur, 1850 m, 13.6.1966, Cheese & Watson 1850 (E); near Ziri, S of Cilo 
Dag, 1830 m, 10.6.1970, Trelawny 1836 (E).
IRAQ: MSU: Oak forest Kopi Qaradagh mt, 29.4.1964, Berkley & Abdul-Rahman 8497 (K); Waziara prope Kopi Qaradagh, 
29.5.1961, Hadad 6282 (MSB); Montes Avroman, 5000', 7.1867, Haussknecht 345 (G-BOIS, M); Penjwin, 1500 m, 
23.4.1947, Rawi 8811 (US); Penjwin, 1400 m, 23.4.1947, Rawi 8946 (MSN); distr. Sulaimaniyeh (Kurdistan), montes 
Avroman ad confines Persiae, in ditione pagi Tawilla, in saxosis calc., 1800-2000 m, 15.-18.6.1957, Rechinger 10343 (B, E, 
M, W); in ditione pagi Penjwin, 1500 m, 19.-20.6.1957, Rechinger 10471 (G, W).
IRAN: PROV. W AZARBAIJAN: Mt Sir, Urmia district, 27.5.1928, Gilliat-Smith 2280 (K). — PROV. ZANJAN: Zanjan-Hamadan 
road, hills above the village Dashti on the lower slopes of Qeydar mt, about 35 km from Soltanieh, 2200-2600 m, 3.7.1974, 
Alava 14268 (TUR); Kuh Anguran, 35-42 km SW Tashvir, inter Manjil et Zanjan, 1900-2200 m, 2.6.1971, Lamond & 
Iranshahr 40885 (W); Manjil to Zanjan, S side of Tarom pass, 42 km from Tashvir, 1900 m, 2.6.1971, Lamond & Iranshahr 
3609 (E); ibid., Iranshahr 14761 (W); Mahneshan, Alam Kandy, Ghar-ghalan mts, 1900-2500 m, 27.6.1983, Moussavi 41070 
(W); Gheydar, 35 km S Soltanieh, Kouhayeh Gheydar, 2200-2650 m, 3.7.1974, Termeh & Moussavi 16457 (W). — PROV. 
KORDESTAN: 70' N of Sanandaj, 6000', 19.5.1962, Furse 2122 (E, K, W); Bijar, Kuh Chang-Almas, 2200-2500 m, 8.7.1968, 
Iranshahr 13252 (W); Salavatabad, 25 km on road from Sanandaj to Hamadan, 2300 m, 3.7.1971, Lamond 4467 (E, M); in 
monte Hamzeh Arab SE Bijar, 2200-2600 m, 1.7.1971, Lamond & Termeh 42594 (M, W); dry rocky slopes above mt pass 26 
km E Sanandaj, c. 2200 m, 28.6.1965, Ledingham & al. 4235 (LE, W); in jugo Ariz 20 km W Sanandaj, 2200 m, 4.7.1971, 
Rechinger 42863 (M, W); in quercetis 90-110 km W Sanandaj versus Marivan, 1650-1800 m, 6.7.1971, Rechinger 42944 (M, 
W); ad versuras 33 km NW Divandarreh versus Saqqez, 2100 m, 29.6.1974, Rechinger 48591 (M, W); Bijar, Kuhe Hamzeh 
Arab, 2000-2600 m, 31.6.1971, Termeh 40931 (W); Sanandaj to Marivan, pass Ariz, 25 km de Sanandaj, 2200-2350 m, 
3.7.1971, Termeh 41022 (W); Sanandaj to Marivan, 80 km after Sanandaj, 1700 m, 5.7.1971, Termeh 40930 (W); between 
Divandarreh and Saqez, 2200 m, 31.5.1974, Wendelbo & al. 11934 (LE, W); 20 km W Sanandaj, inter Sanandaj et Marivan, 
step hillside, c. 7050’, 20.5.1960, Bent & Wright 520-107 (K, W). — PROV. HAMADAN: Assadabad pass, W of Hamadan, 
2220 m, 12.5.1966, Archibald 1807 (E, K); in cacumine montis Elwend, 8.6.1905, Bornmüller 317 (B); Razan to Avaj, 10 
km to Avaj, on top of the neck, 2200-2300 m, 10.7.1994, Chehregani & Zarre 17897 (MSB, TARI, TUH); Abgarm-Avaj, 
road of Mahmud Abad, 1450 m, 14.5.1974, Dini & Bazargan 8676 (W); 13 km SW Hamadan, 2350 m, 8.6.1959, Pabot 1466 
(G, W: under 12430); auf sonnigen Stellen auf den Elwend, bei Haydere, 7000', 6.6.1882, Pichler (W, WU); auf Voralpen 
des Elwend, bei Haydere nächst Hamadan, 15.6.1882, Pichler 217 (G-BOIS); in monte Elwend (media), 1882, Pichler (B, K, 
LE); 40 km SW Hamadan, 9.6.1959, 2300 m, Pabot 1555 (G); 70 miles NE Hamadan, E slopes, 6600', 17.5.1961, Stutz 1292 
(W); in caucamine montis Elwend, 8.6.1905, Strauss (LE); in monte Karagan, 6.7.1906, Strauss (B); Zanjan to Hamadan, 
140 km SE Zanjan, Kouhayeh Takht, 2150-2550 m, 5.7.1974, Termeh & Moussavi 16462 (W); Gardaneh Assadabad, 2150 
m, 9.5.1975, Wendelbo & Assadi 16816 (W). — PROV. KERMANSHAH: in mt, Kuh-i Gawarreh (ad Kerind), 20.5.1909, 
Strauss (B). — PROV. MARKAZI: Sultanabad, 1.6.1967, Klett 12410 (W). — PROV. LORESTAN: Bisheh, 1800 m, 29.5.1937, 
Köie 710 (B, W).
NOT TRACED: Kurdestania Persiae, 1867-1868, Haussknecht (G); [Iran, prov. Hamadan/Kermanshah]: Entre Kermanschah et 
Hamadan, Olivier (P!).


28. Astragalus nevadensis Boiss.
28a. Astragalus nevadensis subsp. nevadensis 

SPAIN: PROV. DE JAEN: Cazorla, Valdeinfierno, 1800 m, Kalk, 11.7.1985, C. Fernández & Cobos (MA); Sierra de Castril, 
1900 m, 7.1903, Reverchon 303 (MSB). — PROV. MALAGA: Sierra de las Nieves, 4.7.1849, Bourgeau 168 (MSB, P); Sierra 
de las Nieves, 12.7.1889, E. Reverchon Pl. d’Andalousie 393 (MA, MSB, P). — PROV. GRANADA: Sierra Nevada, dans la 
région alpine, au Panderon de Velata, 29.7.1851 et 26.7.1851, Bourgeau 1138 (M, MSB); Sierra Nevada, région alpine à la 
Cueva de los Panderones, 23.7.1852, Del Campo 34 (M, MSB); Sierra Nevada, in schistosis, ad 2700 m, 9.8.1923, 
Cuatrecasas 40 (MSB); Sierra Nevada, im Felsschutt zwischen Parador Nacional und Picacho de Veleta, 26.8.1969, B. & W. 
Lippert 10049 (M); Sierra Nevada in pascuis saxosis ad Parderone, 2500 m, 8.1891, Porta & Rigo 470 (M, MSB); ibid., 
1600-2700 m, 3.7.-1.8.1879, Porta & Rigo 134 (M, MSB); Sierra Nevada, 6000-9000', 3.7.-1.8.1871, Porta & Rigo (M); 
Sierra de Gragalemos (bei Nives), 27.6.1906, Reverchon (M); Piacho de Veleta Nevadae, 30.7.1876, Winkler (M); Sierra 
Nevada, Refugios (Granada), 2400 m, 28.7.1967, Zubizarreta 754 (M).


28b. Astragalus nevadensis subsp. muticus (Pau) Zarre & Podlech

SPAIN: PROV. SORIA: Velilla de la Sierra, 1080 m, 27.6.1978, Zubizarreta 16788 (MSB). — PROV. TERUEL: Camarena de la 
Sierro, Pico de Javalambre, 1800 m, 18.7.1981, Castroviejo & Fdez Quirós 6131 and 13214 (M); declives calcáreos del pinar 
de Pinus silvestris, en Valdecerezo, cerca de Alcalá de la Selva, 1600 m, 7.7.1946, Font & Sierra 356 (MA); Javalambre, 
1500-2000 m, 7.7.1895, Pau (M); ibid., 11.7.1983, Peris & al. 5 (MA); in Sierra de Valacloche, 1700 m, 1893, Reverchon 
(M, MA, MSB, P); Alcala de la Selva (E Teruel), 1500 m, 18.6.1976, Zubizarreta 13699 (M, MA); Camarena, 1700 m, 
4.7.1975, Zubizarreta 8133 (M). — PROV. VALENCIA: Sierra de El Toro, 26.7.1886, Pau (MA); Llanos de El Toro, 1000 m, 
19.6.1919, Sennen 3873 (MSB).


29. Astraglus noeanus Boiss.

TURKEY: B5 YOZGAT: Ankara-Yozgat, 2 km W Yozgat, 1300 m, 7.7.1983, Nydegger 18033 (G); 12 km N Sorgun, 1080 m, 
26.6.1987, Nydegger 42938 (MSB); in declivibus siccis 17 km ad occidentem pagi Akdagmadeni versus Sorgun, 1200-1300 
m, 5.7.1977, Rechinger 57585 (M, W); E Yozgat, 1150 m, 27.6.1970, Sorger 70-1510 (E). — B6 SIVAS: 37 km SW on 
Kayseri road, 4500', 16.6.1962, Apold & al. 30 (E); Hafik, Topaktas area, 1250 m, 27.6.1991, Aydoglü & al. 3457 (GAZI); 
Hafik, near Arpayau village, 1480 m, 1.6.1991, Aydoglü & Hamzaoglü EH396 (GAZI); Pontus australis, Siwas, in apricis, 
1300-1400 m, 9.6.1890, Bornmüller 2126 (B); Yozgat-Sivas, 14 km W Yildizeli, 1490 m, 8.7.1983, Nydegger 18045 (G); 
Kayseri-Sivas, 2 km W Kayadibi, 1340 m, 13.6.1992, Nydegger 46249 (MSB); inter Sivas et Yozgat, 47 km WSW Yildizeli 
ad viam versus Akdagmadeni, 8.9.1957, Rechinger 15240 (M, W); Vilayet Sivas, in declivibus stepposis ad oppidum, 
17.6.1939, Skrivanek (PRC). — B7 ELAZIG: Charput de Sivas, 19.7.1891, Girard de Césaráe (G).


30. Astragalus ochrochlorus Boiss. & Hohen.

IRAN: PROV. MAZANDERAN: In reg. alp. m. Demawend, c. 3000 m, 12.7.1902. J. & A. Bornmüller 6890 (B); Elburs occid., 
supra Iunesar in jugo inter v. Lar et Dschadjerud, 3000 m, 12.7.1902. J. & A. Bornmüller 6891 (B, BM, P, W); Demawend, S 
slopes, 7000', 12.6.1962, Furse 2534 (E, K, W); Demawend, Pelur, 2700 m, 21.7.1935, Gauba 473 (B); Demawend ober 
Rehne, SW-Hang einer Schlucht, 3300 m, 25.7.1936, Gilli (W); E slope of Demavend, 7 km SW Reyneh, 4000 m, 29.7.1964, 
Grant 16539 (W: mixed with A. iodotropis); ibid., 6 km SW Reyneh, 3200 m, Grant 128 (W); Kuh-i Kakashan, 10000', 
18.7.1940, Koelz 16363 (E, MSB, W); E of Damavand, Riene, c. 85 km E Tehran, 8000’, 22.6.1965, Ledingham 4174 (LE, 
W); Damavand, Daryache-ye Tar, Kuh-e Miandarya, 2800-3300 m, 8.6.1980, Moussavi & al. 39869 (W); Larijan, Rineh, 
Kuh-e Damavand, 2750 m, 20.7.1980, Moussavi & al. 40987 (W); Larijan, Rineh, Gousefand Sara, 12 km NW Rineh, Kuh-e 
Damavand, 3000-4200 m, 22.6.1982, Moussavi & al. 41124 (W); 10.5 km N Plour, 2380 m, 10.7.1960, Pabot 4172 (G); 
Parvar Protected Region, in montibus inter Shahmirzad et Fulad Mahalla, 68 km NE Semnan, in declivibus saxosis, 2200 m, 
30.5.1975, Rechinger 52334 (M, W); Pentes S du Kuh-i Demavend, 2000-3500 m, 18.-20.7.1956, Schmid 6415 (G, W); 
Demavend area near Pulur, 2400 m, 22.6.1959, Wendelbo 2065 (BG, W). — PROV. SEMNAN: Zentral Elburs, im Gebiet des 
Kuh-i Nizwa, Berg c. 8 km N Djaschm (Tschafte), 2900 m, 29.7.1948, Behboudi & Aellen 1041 (W); Zentral Elburs, 
Oberlauf des Tscheschme-i Ali, SO des Kuh-i Nizwa, Bergsteppe und Felsschutt N Djaschm (Tschafte), 2200-2600 m, 
28.7.1948, Behboudi & Aellen 1045 (W); Elborz, Kuh Nizva, 29.7.1948, Behboudi & Aellen 5591 (W); in declivibus 
australibus montium Shavar ad supra Nekarman, 2300 m, 20.-26.7.1948, Rechinger 5842 (E, W); Nezva Kuh area, S of and 
above Taru, c. 2400 m, 7.7.1959, Wendelbo 1210 (BG, W). — PROV. TEHRAN: Damavand, 2400 m, 16.8.1973, Amin 15128 
(W); Zentral-Elburs, Flussebene des oberen Tedschen bei Kom-rud pain, 60 km östlich von Firuzkuh, c. 2200 m, 24.7.1948, 
Behboudi & Aellen 1025 (W); rocky S foothills of Elburz mts. Ab Ali, c. 60 km E Tehran, 2.6.1965, Ledingham 4170 (LE, 
W).
NOT TRACED: Sefid Khok, 2400 m, 2.-3.6.1956, Schmid 5933 (G, M, W), 5935 (W) and 5969 (G).

Specimens with well-developed peduncles:
PROV. MAZANDARAN: E Tehran, Lar, Kharsang, 2500 m, 13.8.1973, Amin 15365 (W); Lar valley, 2450-2550 m, 2.7.1974, 
Wendelbo & Assadi 13368 (LE, W).


31. Astragalus oltensis Grossh.

TURKEY: A6 TOKAT: Yildizeli to Akdagmadeni, Seyh Halil Tikkesi village, 1150 m, 9.9.1994, Adigüzel & Aytaç 1848 
(GAZI); A7 GÜMÜSSANE: Sipikuri to Köse, 26.6.1934, Balls 1551 (E).


32. Astragalus oreites Beck
Only known from the type gathering.


33. Astragalus polyanthus Bunge
33a. Astragalus polyanthus subsp. polyanthus

IRAN: PROV. W AZARBAIJAN: 15 km SW Shavat towards Siah-Cheshmeh, 1500-1650 m, 24.6.1991, Akhani 7732 (MSB); 
Meshow dagh, 30' NW Khoi, 6000', 3.8.1963, Furse 3534 (K); Ghoje Dagh, near Bazargan, c. 2000-2250 m, 1.8.1971, 
Lamond 5005 (M); in jugo 30 km NW Khovy, 1700-1800 m, 2.8.1965, Rechinger 32695 (M, W); in monte Ghojeh Dagh, W 
Bazargan ad confines Turciae, 2100-2250 m, 1.8.1971, W. Rechinger 44003 (W); in declivibus saxosis inter vallem fluvii 
Rud-e Aland et pagum Qeris, NW Khovy, 2100 m, 16.7.1974, Rechinger 49489 (W).


33b. Astragalus polyanthus subsp. vedicus (Takht.) Zarre

ARMENIA: Araratskyi khrebet, 27.6.1970, Tapakuap (E); Vediskyi raion, Boz-Buron, 800 m, 30.6.1948, Akhverdov & 
Mirzova (E); Daralagez, 2000 m, 21.7.1950, Akhvergov & Mirzova (E).


34. Astragalus prominens (Boriss.) Boriss.
Known only from the type gathering.


35. Astragalus sahendi Fisch.

IRAN: PROV. E AZARBAIJAN: Shahbil, Kuh-e Sabalan, 3470 m, 9.7.1974, Foroughi & Assadi 13941 (E, W); Sähend prope 
Tabris, 6.1859, Bunge & Bienert (K: photo MSB, G: photo M); Ispereichan, in montis Sahend elatoribuls, 4.8.1884, Knapp 
(B, JE, WU); Kuhhaye Sabalan, 3500-3700 m, 15.6.1971, Lamond 4770 (E); Montes Sabalan, in declivibus borealibus 
saxosis, 3500-3700 m, 15.7.1971, Lamond & Termeh 4770 (in Rechinger 44159) (M, MSB, W); ibid., 44158 (W); ad fines 
rossicas, mons Sawellan, 5.1898, Lewandowsky 1714 (C, G: photo M); in alpe Sahend, 5.-17.8.1856, Seidlitz (G: photo M 
and MSB, P); Sardab to Kuh Sabalan, 24.7.1968, Termeh 13264 (W); Ardabil, Ghotour-Sou, Kuh-e Sabalan, 3400-3600 m, 
15.7.1971, Termeh 40820 (W); Bostan-abad, Pishiclou to Chiniboulagh, 4 km from Pishiclou, 2150-2350 m, 1.8.1984, 
Termeh & Moussavi 41386 (W); Oskou to Sahand, Kuh-e Soltan, 2600-3130 m, 22.6.1985, Termeh & al. 41589 (W).


36. Astragalus sangesuricus Boriss.

ARMENIA: Megri distr., 1700-2200 m, 14.7.1968, Mulkijanian & al. (E).


37. Astragalus sempervirens Lam.

SWITZERLAND: Valle Findelin, Zermatt, 2200 m, 8.11.1903, Spencer (M).
FRANCE: DEPT. BASSES-PYRÉNÉES: Col d’Arnousse, 5.8.1893, Foucaud & Gamin 3447 (MSB, P). — DEPT. HAUTES 
PYRENÉES: Vallée d’Héas, 7.8.1903, Sennen (MSB). — DEPT. PYRÉNÉES-ORIENTALES: Gavarnie, Vallée de Pouey d’Aspé 
(M). — DEPT. HAUTES-ALPES: St.-Firmin, le Grün, les Reculaz en Valgodemar, 1000-1500 m, 15.5.-15.7.1882, Darnaud 
3676 (MSB, P); à La Grave, 1800 m, 20.7.1858, Mathonnet in el. Mailleanae 640 (MSB, P). — DEPT. ALPES-DE-HAUTE-
PROVENCE: Castellan, am Verdon Fluß, 2.8.1962, E. Berger (M); Aurent, près Annot, 18.7.1887, Reverchon & Derbez in Pl. 
Franc. 191 (MA); Allos, vallion de Méouille, 2100 m, 27.7.1984, de Retz 85533 in Soc. Ech. Pl. Vasc. 12248 (M, MSB). — 
DEPT. ALPES-MARITIMES: Col de Tende, 17.7.1861, Bourgeau Pl. Alp. mar. 85 (MSB, P); envir. de Beuil, 1500-1600 m, 
7.1894, G.Vidal in Fl. select. exs. Magnier 3496 (BC, MSB, P).
SPAIN: PROV. LEÓN: Maraña, Mampodre, Pico Salamanquino, 1520 m, 27.7.1986, Urrutia & al. 1457.86 (MA, MSB). — 
PROV. SANTANDER: Picos de Europa, 8.1945, Marten (BC, MSB).
ITALY: PROV. CUNEO: der Cma. d. Becco O des Colle di Tenda, 2100 m, 6.8.1972, Podlech 24064 (M, MSB). — PROV. 
IMPERIA: Mte Grai N Pigna, 1750-1900 m, 30.7.1984, Lippert & Merxmüller 20121 (M). — ALPI MARITTIME: Limone 
Kalkhänge und Matten über Maria Cross, 1400-1800 m, 28.7.1951, Merxmüller & Wiedmann 10755 (M); entre Colla Melsa 
et Cima di Marta, 1600-1800 m, 23.6.1976, Moggi & Ricceri 8383 (M). — PROV. L’AQUILA: Am Passo Capannello, N 
L’Aquila, c. 1300 m, 18.8.1965, Merxmüller & Grau 20682 (M). — PROV. NÁPOL: In pascuis suprá Piano cinque Miglia, 
Apratii, 18.6.1856, Huet Du Pavillon 306 (M). — PROV. LECCE: Apuanische Alpen, von Gramolazzo zum Rifugio Donegani, 
700-1250 m, 19.5.1965, Merxmüller & Wiedmann 19629 (M). — PROV. POTENZA: M. Seriu, in pascuis aridids del Papa, 
1900-2000 m, 26.7.1898, Dörfler 491 (M).
GREECE: PROV. LEFKÁDA: In pascuis montis Ainos ad cacumen in insula Cephalonia, 7.1879, Letourneux 3372 (W). — 
PROV. KEFALONIÁ: In monte Nero Cephalonicae, 3000 pedes, 13.10.1834, Schimper & Wiest. (W). — PROV. FOKIDA: In m. 
Oeta Phthiotidis, in pascuis subalpinis, 5500', 12.7.1879, Heldreich (W). — PROV. VIOTÍA: In monte Parnassi, 8.1852, 
Heldreich 2674 (LE, W); in regione supriori montis Parnassi, 3000-7000', 15.7.-27.7.1854, Orphanides 385 (W). — KRÍTI: In 
mt. Creta, 1879, Heldreich (W).


38. Astragalus titziae Ghahreman & Zarre

Iran: PROV.TEHRAN: 20' SE Firuzkuh, 9000', 1.7.1962, Furse 3010 (K); Seyyedabad, road of Firuzkuh, 2500 m, 4.6.1972, 
Dini & Arazm 15656 (TARI, W); Firuzkuh to Tehran, 28 km from Firuzkuh, Kuh-e Velveleh, 1950-2200 m, 28.5.1980, 
Termeh & al. 39850 (W).


39. Astragalus transoxanus Bunge

UZBEKISTAN: Sewerni Makrosklon, khr. Koitash, Tsentralnoi usadbi sowkhoza, im. Kirova, Kamenistie sklon, 19.6.1971, 
Botchansev & Kamelin (LE); Samarkand, okr. Dzhizaka, Swerni sklon Gori Nadi Irane-Kishlakom, 11.4.1915, O.A. & B. 
Fedtschenko 196 (LE); Samarkand, Dzhizakskii kishd Bagadani, 25.5.1914, Knorring (LE); ad decliva saxosa montium 
Kara-tau (in adjacentibus st. v. f. Kuropatkino) prope pag. Sarybulak, 600 m, 9.6.1929, Kurdjaschev (in hb. Fl. SSR 1413 or 
6271: E, G, MSB, P, W); Nuratinskyi Khrebet, Ak-mau, inter Tumshuk et Lyangar, 5.-7.5.1951, Neuchreva 64 (MSB).
TADZHIKISTAN: Prope Samarkand, in apricis ad fl. Zeravschan versus, 2200', 16.6.1901, Alexeenko 132 (LE); Turkestan, 
Ouromitane, 15.6.1881, Capus 351 (MSB, P); Sarewshan Gebiet, Pendschakent, 3000', 17.-29.6.1882, Regel (MSB).


40. Astragalus yueksekovae V. A. Matthews

TURKEY: C10 HAKKARI: 2150 m, 26.6.1966, Davis 45653 (E); ibid., 2000-2100 m, 30.6.1966, Davis 45820 (E); Sat Dagi, 
first ridge between Yüksekova & Varegöz, Aruna Geçidi , c. 2500 m, 24.7.1983, Ekim 8010 (GAZI); ad pontem Yeni Köprü 
inter Yüksekova et Shivelan, 1800 m, 30.6.1975, Rechinger 53848 (M, W).
IRAN: PROV. W AZARBAIJAN: Oroumieh, Razhan, Khalil Kuh, 1600-1800 m, 9.7.1994, Chehregani & Zarre 17873 (MSB, 
TUH); in monte Chalil Kuh prope Razhan, 2600-3200 m, 2.7.1974, Rechinger & Renz 48765 (M, MSB, W); ibid., 1800-2000 
m, 2.7.1974, Rechinger 48754 (M, W).



Astragalus sect. Macrophyllium Boiss.

1. Astragalus cephalotes Banks & Sol.

TURKEY: A5 ÇORUM: Before Osmancik, 600 m, 31.5.1969, Tobey 2655 (E). — A8 ÇORUH (ARTVIN): Çoruh gorge, between 
Artvin & Ardanuc, 26.6.1957, Davis & Hedge D.30054 (E); A4 ANKARA: Natolia, prope Angora (Ankara) in m. Dikman, 
1834, Wiedemann 7 (G-BOIS, LE). — A4 KIRIKALE: Delice, Bayakavazar kasabsi, c. 1150 m, 26.6.1991, Yaman 1015 
(GAZI). — B5 YOZGAT: E Ankara, zwischen Ankara und Sivas, c. 20 km SE Sungurlu, Ruinengelände von Hattusas bei 
Bogazkale, 17.6.1987, Lange 3543 (Hb. Lange). — B5 KAYSERI: Ad pagum Tschomakli, 29.5.1903, Bornmüller (L). — B6 
KAYSERI: Sariz, Yolak (Yesilkent) Dayoluk köyü, Koca Dag teppesi, 1750 m, 5.8.1988, Aytaç 2513 (GAZI). — B6 
MALATAYA: Kavak Aghateh [Akça Dag] to Arga, 16.7.1906, Post 134 (G); 50 km N Malataya, Pazarcik, 17.5.1975, Browicz 
& Zielinski 250 (E); on rocky hillside near the village of Kozluk, in the upper course of the river above Izmet Pasa, 8.6.1968, 
Alava 7074 (E). — B7 ELAZIG: Tunceli to Elazig, zw. Munzur Suyu und Peri Suyu, 1050 m, 25.8.1987, Engel 170 (MSB). — 
Elazig to Pertek, 1300 m, 6.6.1957, Davis & Hedge 29197 (E, M). — Elazig-Kale, (on Euphrates), 22 miles E of Elazig, 
1300 m, 4.6.1957, Davis & Hedge 28940A (E). — C5 IÇEL: S foot of Bolkardaglari, between Yavca and Arslanköy, NE of 
Mersin (Içel), 1.6.1991, Boratynski & al. 6856 (MSB); Mersin (Içel), Delikkaya - Cambarli, 810 m, 26.6.1972, Uslu 1550 
(E). — C5 ADANA: Pozanti, 3 miles N on E slopes of Toros Dagi, 1000 m, 13.7.1971, Abb. Uni. Aman. Exp. B170 (E); 3-4 
km N of Pozanti, 1200-1700 m, 31.7.1971, coll. ignatus NI 799 (E); Karinca Da., 2 miles NE Alpu, 3 miles N Pozanti, 1100', 
18.7.1971, Abb. Uni. Aman. Exp. F2 428 (E); Pozanti-Tekir Yayla, Pinus brutia forest, Vural 6866 (GAZI); C6 ADANA: 
Osmanyie, Yarpuz village, Soku area, 1500 m, 20.7.1988, Aytaç 2439 (GAZI). — C6 KAHRAMANMARAS#S: Ahir Dag above 
Kahramanmaras#, 1100 m, 2.5.1957, Davis & Hedge 27472 (E); Vil. Maras, Valley of Tekir river near Sucati, NE Alikayasi, 
stony places, 6.6.1991, Boratynski & al. 6887 (MSB); Öksüz Dagi, Tarla kenari, 1400-1500 m, 14.6.1987, Duman 3277 
(GAZI); Öksüz Dagi, 1500-1600 m, 26.7.1987, Duman 3766 (GAZI); Engizek Dagi, Aksu mahallesi çevresi, 1000-1100 m, 
24.5.1987, Duman 2888 (GAZI); above Marash, 2700', 2.5.1934, Balls 954 (E). — C6 GAZIANTEP: Alep [Aleppo, Syria] – 
Aintab [Gaziantep], 20.5.1937, Férer Louis 94 (MSB); W of Gaziantep, 19.5.1972, Rückbrodt (E). — C7 URFA: Urfa-Hilvan, 
15 km from Urfa, 18.5.1957, Davis & Hedge D.28255 (E); in apris c. Orfa [Urfa], 1600 ped., 15.5.1865, Haussknecht [45] 
(G-BOIS, P, W). — B8 DIYARBAKIR: Diyarbakir-Ergani, 10 km from Diyarbakir, 750 m, 1.6.1957, Davis & Hedge D.28798 
(E); 69 km N Diyarbakir, on Diyarbakir to Lice road, ± 800 m, 10.5.1988, Cribb (K). — C8 MARDIN: Mardin, Senar, 
6.7.1888, Sintenis 930 (BR, G, GOET, P, WU).
LEBANON: Rascheya, 5.1846, Boissier (G, G-BOIS); ad Antilibani radices occidentales, in declivitatibus supra Baalbeck, 
1150-1300 m, 19.5.1910, Bornmüller 11586 (B); Wadi Yenayli above Baalbak, 5000', 23.6.1943, Davis 6518A (E); 
Baalbeck, 24.8.1930, Gombault 1373 and 14.5.1933, Gombault 2051 (MSB, P); a Zachle, 1.7.1930, Gombault 1371 (P); 
Antiliban, Ouadi Rheiri, 1600 m, 21.7.1951, Pabot (G); Baalbeck, 27.5.1893, Peyron (P); Zachle, 20.7.1914, Pastuchov 
(LE).
SYRIA: Pr. Damasc, in campis derelictis versus Palmyram, 4500', 26.5.1855, Kotschy 589 (G-BOIS, P); Kurdistania 
occidentalis, in altiplanitie ad meridiem vici Kjachta districtus Mamuret-ül-Asis ad Euphratem versus, in declivibus lapidosis 
infra viam Parwadjak, 700 m, 10.7.1910, Handel-Mazzetti 407 (1966) (W, WU).
IRAQ: MAM: Mosul liwa, Bakrima, 1300 m, 9.5.1947, Rawi 8966a (US); distr. Mosul (Kurdistan), ad confines Turciae prov. 
Hakkari, in ditione pagi Sharanish, in montibus calc. a Zakho septentrionem versus, Jabal Khantur, 1200 m, 4.-9.7.1957, 
Rechinger 10742 (W); distr. Mosul (Kurdistan), ad confines Turciae prov. Hakkari, inter Dohuk et Amadiya supra Smara 
Tuka, 1500 m, 10.-12.7.1957, Rechinger 11533 (W); ibid., 1200 m, Rechinger 11642 (W); distr. Mosul (Kurdistan), ad 
confines Turciae prov. Hakkari, in ditione oppidi Zakho, in faucibus c. 5 km a Zakho meridiem versus, 2.-4.7.1957, 
Rechinger 10717 (M, W). — MRO: Shaqlawa, 900 m, 8.5.1947, Gillett 8062 (K); Pir Omar Gudrun, 5000', 6.1867, 
Haussknecht 343 (BM, G-BOIS, LE, P, W). — MSU: inter Sulaimanyia et Qaradagh, loco Dar Mazala dicto, 28.5.1961, 
Hadad & al. 5105 (MSB); Darband-i Basian, 1867, Haussknecht [339](G-BOIS, P); inter Sulaimaniya et Dokan, in jugo 
prope Surdash, in apertis Quercetorum, c. 1000 m, 14.6.1957, Rechinger 10117 (W). — FKI: in collibus siccis 37 km a 
Kirkuk orientem versus, c. 600 m, 13.6.1957, Rechinger 10043 (B, M, W).
IRAN: PROV. W AZARBAIJAN: In declivibus australibus jugi Gardaneh-ye Zamziran S Mahabad versus Sardasht, 1200 m, 
8.7.1974, Rechinger 49095 (M, W).
NOT TRACED: Kurdistania, Süverek, Kera Kaei, 23.5.1888, Sintenis 763 (G, P); [Turkey] Cappadocia, Baphry, Aucher-Aloy 
1250 (P); [Turkey] Cilicien, Borakali, 800 m, 6.1895, Siehe 183 (LE); [Turkey] Kalkige Abhänge Tarbas, 1300 m, 20.6.1898, 
Siehe 139 (LE); inter Orul et Nirib, 23.6.1865, Haussknecht (P); Coelsyria, Hammara, 17.7.1932, Gombault 1373 (P); 
Barouk, 1.7.1937, Frér Louis (P); Syria/Lebanon: Entre Ain menelouwre et Onadi et Karn, Antiliban, route de Damascus à 
Bayrouth, 15.5.1856, Gaillardot 1761 (G-BOIS); plaine entre Antiliban et Damas, 5.-6.1846, Boissier (G-BOISS).

Specimens with a short calyx as in the type of A. andrachnaefolius:
TURKEY: A4 KASTAMONU: Bölükyazi köyü, Tasköprü, 4.6.1967, Baytop 11338 (E) — A8 ÇORUM: 6 km W Altiparmak NW 
Yusufeli, 18.6.1988, 1780 m, Nydegger 43390 (MSB). — A5 AMASYA: Ad Turkhall inter Amasia et Tokal, 600-700 m, 
29.5.1890, Bornmüller 2124 (B) – Pontus Galaticus, Kara dagh, inter Amasia et Samsoun, 3.6.1889, Bornmüller 88 (B, WU). 
— C5 IÇEL: Région montagenuse du Taurus, pres du défilé des Portes Ciliciennes [Gülek Bogazi], 23.7.1855, Balansa 485 
(G, MSB, P); Gülek-Bogazi, 1300 m, 20.6.1987, Nydegger 42877 (MSB). — C6 GAZIANTEP: Aintab, in agris, 20.6.1865, 
Haussknecht [44] (G-BOIS, LE, P); in apris calc. c. Aintab, 2000 ped., 20.6.1865, Haussknecht (W).
SYRIA: Aleppo, Boissier (G-BOIS); Muslimiya, 4.4.1936, Frér Louis 94 (P).
IRAQ: MRO: Kurdish hills near Zawita, 3000', 1.5.1936, Mrs Low 271 (BM).


2. Astragalus dipodurus Bunge

TURKEY: C4 KONYA: Armusun village near Karasiner, 75 km S of Konya, c. 1100 m, 15.7.1965, Ledingham & al. 4351 (E, 
LE); 24 km S Seydiss#ehir, 1050 m, 30.7.1992, Nydegger 46898 (MSB). — C6 KAHRAMANMARAS#s: Çaglayancerit - Bozlar, 
c. 1000 m, 2.6.1987, H. Duman 3103 (GAZI); Çaglayancerit, Öksüz Dagi, 1300-1600 m, 6.7.1991, Aytaç & Duman 4065 
(GAZI); Beryt Dagh, 1865, Haussknecht (G-BOIS); 15 km to Çaglayancerit from Pazarcik, open Quercus forest, 580 m, 
14.8.1997, Zarre 157 (E, MSB); Çaglayancerit - Bozlar, 1000 m, 14.8.1997, Zarre 173 (MSB). — C6 URFA: Birecik-Halfeti, 
8 km S Halfeti, 600 m, 27.5.1987, Nydegger 42523 (MSB).
IRAN: PROV. W AZARBAIJAN: In declivibus borealibus jugi Qushchi inter Shahpur et Rezaiyeh, 1700 m, 21.7.1974, Rechinger 
49807 (M, W).


3. Astragalus glabrifolius Bunge

AFGHANISTAN: PROV. GHAZNI: Tangi, an der Straße von Malestan nach Dasht-i-Nawr, 2370 m, 13.8.1970, Podlech 19297 
(M, MSB); Wulghan, an der Straße von Malestan nach Sange Masha, 2680 m, 14.8.1970, Podlech 19378 (G, M, MSB); 2 km 
W Dahangol, an der Straße von Malestan nach Namut, 2780 m, Silikat, 30.6.1978, Podlech 31972 (G, MSB); distr. Malestan, 
inter Miradina et jugum Ghoutch Kol, N Sang-i Masha, 3300 m, 2.7.1962, Rechinger 17603 (M, W); Sariab 52 km W 
Ghazni, 17.7.1967, Rechinger 37166 (W). — PROV. KABUL: Kabul, Topdara, 5.9.1924, Bukinich (LE); bei Kabul, beim Tob 
am Scher Darwasah, 1840 m, 11.6.1951, Gilli 1533 (W); bei Kabul unter der Grabmoschee Safa am Scher Darwasah, 1900 
m, 27.6.1949, Gilli 1534 (W); bei Kabul, Berg W des Asmani unter dem Gipfel, 1920 m, 3.8.1949, Gilli 1535 (W); Paghman, 
9000', 27.6.1937, Koelz 12137 (W); Istalif, road between main road and village, 1900-2000 m, 22.6.1965, Lamond 2009 (E, 
M, W); Istalif, 1900 m, 22.6.1965, Rechinger 31197 (M, W); in declivibus montium 10 km NW Kabul ad viam versus 
Charikar ducentem, 2000-2100 m, 21.6.1965, Rechinger 31168 (W). — PROV. PARWAN: Mittleres Salang-Tal bei Qalatak, 
1950 m, 6.10.1970, Podlech 19713 (M, MSB). — PROV. BAMIAN: Nawa-i-Waras 1 km oberhalb der Einmündung in Nawa-i-
Surkhjoy an der Straße Panjaw-Sharestan, 2730 m, 28.7.1970, Podlech 18908 (M, MSB); 20 km SW Panjao versus jugum 
Waras, 2500 m, 30.6.1967, Rechinger 36540 (M). — PROV. OROZGAN: Südhang des Kotale Karkarake Naw an der Straße 
nach Daykundi, 2280 m, 30.7.1970, Podlech 19060 (M, MSB, W); 7 km O Orozgan, 2250 m, 29.6.1978, Podlech 31898 (G, 
M, MSB); Syahdara bei Dahane Khubabak an der Straße nach Daykundi, 2420 m, 29.7.1970, Podlech 19031 (G, M, MSB); 
Deh Kundi, in saxosis et arenosis graniticis 3-30 km NE Sharestan, 2200 m, 1.-2.7.1967, Rechinger 36765 (W).
NOT TRACED: Isphi, 8000', 7.6.1937, Koelz 11755 (E: photo MSB, W); Hazarajat, S side of Kuh-i-Baba, near Sultan Ribat, 
8500', 29.8.1954, Thesiger 116 (W).


4. Astragalus isauricus Hub.-Mor. & V.A. Matthews

TURKEY: C3 KONYA: Seydisehir - Akseki yolu, 15 km, 1665 m, 24.7.1997, Duran 4238 (GAZI, MSB). — C4 KONYA: 
Ermenek, Damlacali, 1750 m, 7.7.1978, forest of Cedrus libani, Vural 960 (E, GAZI).


5. Astragalus karjaginii (Boriss.) Boriss.

AZERBAIJAN: NAKHICHEVAN: Shachbuz region, Kizil-bpez, 900 m, 31.7.1947, Grossheim & al. (BM); Schachbuzskyii raion, 
Kizil-bpaz, 900 m, 31.5.1947, Grossheim & al. (LE); ad portam Darlagez in lapidosis siccis, 14.8.1934, Prilipko (LE); Pr. 
pag. Bliz, S Surab, Nachitchevan, 16.7.19??, coll. ignatus (LE: pp. A. compactus).
ARMENIA: Distr. Ararat, montes Gegamski khrebet, loco Aruni Dzor dicto, 1500-1900 m, 11.7.1975, Vašák (B, G, M); Distr. 
Ararat, montes Gegamski khrebet, loco Ganly Dara dicto, in declivibus occidentalibus fluminis Khosrov, 1400-1600 m, 
13.10.1974, Vašák (G, M); Vedi region, 1750 m, 14.7.1948, Akhverdov (E); Sklon Zangezurskogo khrebta, 1.7.1952, An. 
Fedorov (LE).
IRAN: PROV. W AZARBAIJAN: Oroumieh, Razhan, Khalil Kuh, 9.7.1994, Chehregani & Zarre 17878 (MSB, TUH); Shahpur 
to Rezaieh, Ghoushchi, 1600 m, 12.6.1971, Iranshahr 40897 (W); Qushchi pass between Shahpur and Rezaiyeh, N side, 
1800 m, 12.6.1971, Lamond 4093 (E); 26 km W Rezaiyeh towards Serow, steep loose conglomerate rocks, 1600-1700 m, 
14.6.1971, Lamond 4132 (E); in jugo Qushchi inter Shahpur et Rezaiyeh, 1600-1850 m, 13.6.1971, Rechinger 41883 (M, W); 
a Rezaiyeh 26 km NW versus Sero, 1600-1700 m, 14.6.1971, Rechinger 42021 (W); Chalil Kuh, in faucibus NW Selvana, 
1750-2000 m, 4.7.1974, Rechinger 48932 (M, W).


6. Astragalus longifolius Lam.

TURKEY: A8 ERZURUM: Erzurum - Tortum (Richtung Artvin), c. 25 km N Erzurum, 1750 m, 18.8.1987, Engel 145 (MSB); 
Erzurum - Ispir (Rize), vor Ispir, 2150 m, 19.8.1987, Engel 151 (MSB). — B7 ELAZIG#: Elazig# - Bingöl arasi, Hosmat 
Karayollari Park, korunmus saha, c. 1200 m, 21.6.1983, Ekim 7310 (ANK, GAZI). — B9 BITLIS: Nemrud Dag, im Krater am 
Nemrud Gölü, 2300 m, 9.8.1987, Engel 102 (MSB); E flank of Nemrut Dag, 6000', 3.7.1954, Polunin D.23546 (E); Tütün 
Bakimevi, c. 1500-1600 m, 23.6.1983, Ekim 7517 (GAZI).
IRAQ: MAM: Distr. Mosul, ad confines Turciae prov. Hakkari, inter Dohuk et Amadiya, in quercetis saxosis supra Sirsank, 
1200 m, 10.-12.7.1957, Rechinger 11699 (E, M, W).
NOT TRACED: Armenia orientalis, “Tragacantha orientalis, foliis siccis glabris....”, Tournefort cor. 3 (G-DC).


7. Astragalus octopus C.C. Towns.

IRAQ: MSU: Distr. Sulaimanya (Kurdistan): Montes Avroman ad confines Persiae, in ditione pagi Tawilla, 1800 m, 15.-
18.6.1957, Rechinger 10345 (W); ibid., c. 2000 m, Rechinger 10379 (W).
IRAN: PROV. KORDESTAN: 75 km WNW Sanandaj toward Marivan, 1890 m, 18.5.1966, Archibald 2041 (E, K, W); c. 130 km 
from Sanandaj on road to Marivan, W side of pass, c. 1600 m, 6.7.1971, Lamond 4558 (E); in quercetis (Qu. brantii) 90-110 
km W Sanandaj versus Marivan, 1650-1800 m, 6.7.1971, Rechinger 42916 (W); Montes Chehel Chashmeh, 44 km NE 
Marivan, 2000 m, 7.7.1971, Rechinger 43069 (W); Sanandaj to Marivan, 110 km to Sanandaj, 1600 m, 5.7.1971, Termeh 
40913 (W); Marivan to Saghez, 90 km to Marivan, 1900 m, 7.7.1971, Termeh 40880 (W).


8. Astragalus oleaefolius DC.

ARMENIA: Distr. Ararat, montes Gegamskyi khrebet, loco Aroni Dzor dicto, 1500-1900 m, 11.7.1975, Vašák (M); Distr. 
Ashtarak, in declivibus montis Arailer in vicinitate pagi Egvart, 1300-1900 m, 15.7.1975, Vašák (M); distr. Mikojan, 
Chatschik, 22.7.1950, Ivanova (M).
AZERBAIJAN: NAKHICHEVAN: Prope p. Bist in lapidosis, 19.6.1933, Prilipko (LE); ibid., 24.8.1931, Prilipko & Vichert (LE).
TURKEY: A7 GÜMÜSANE: Szandschak Gümüschkhane, Nuvena-deressi, ad versuras prop. Baijana-Kala, 19.7.1894, Sintenis 
7142 (B, M, G, LE, MSB, P, WU). — A8 BAIBURT: In decliv. aridis montis Berdak, pr. Baibout, 28.6.1862, Bourgeau 77 (P). 
— B8 ERZURUM: Erzurum - Ispir, 50 km after Erzurum, Egerti köyü kuzeyi, Kelkaya mevkii, 2200 m, 29.7.1989, Aytaç 2949 
(GAZI); ibid., 19.7.1990, Duman & Aytaç 3205 (GAZI); ibid., 11.8.1988, Aytaç 2561 (GAZI). — B2 AFYON: Afyon - 
Kütahya yolu, 20 km after Afyon, 1000 m, 3.8.1993, Aytaç (GAZI). — B3 AFYON: Sandikli - Dinar arasi, c. 10-15 km , 1000 
m, 21.8.1993, Duman & al. 5253 (GAZI). — A4 ANKARA: Galatia, in Agro Ancyritano, ad montem Dikmen-dagh, valle 
Kawakli-dere, 900-1000 m, 2.6.1929, J. & F. Bornmüller 13997 (B); Cubuk baraji, 13.7.1955, H. & E. Walter 1331 (B); 10 
km N of Ankara, vineyard on devastated Quercetum, 13.7.1962, M. & D. Zohary 337 (MSB). — B4 ANKARA: Haymana 
Karacaören K., Demirhavan Tepe, 1040 m, 26.9.1992, Vural & al. 6212 (GAZI). — B5 NEVSEHIR: Göreme, Elnazar Kilisesi 
civari, 1140 m, 17.6.1989, Vural & al. 5150 (GAZI). — B5 KAYSERI: Argaeus, Erdschias-dagh, auf steinigen Abhängen des 
Ali-dagh, c. 1700 m, 6.1902, Zederbauer (WU). — B6 SIVAS: Sarkisla-Ortaköy, 1250 m, 27.8.1994, Aytaç 6718 (GAZI). — 
B7 ERZINCAN: Egin, Kyl-Maghala-dagh, 5.6.1890, Sintenis 2878 (B, G, P, WU). — B7 ELAZIG#: Elazig#-Bingöl arasi, 
Hosmat Karayollari Park, korunmus saha, c. 1600 m, 21.6.1983, Ekim 7310 (GAZI). — B8 MUS#S: Mushskii sandzhak, 
Bostankaya, skali, 29.6.1916, Shishkin (LE). — B9 AG#RI: Ercis-Agri, 20 km vor Patnos bei Karabasan (7–8 km v. d. 
Hauptstr.), 2200 m, 15.8.1987, Engel 133 (MSB). — B9 VAN: prov. Hakkari, Zap Gorge, 30 miles S Baskale, 2.8.1954, Davis 
23820 (ANK, E); van to Edremit, Doganli köyü mezarlik, 1750 m, 24.8.1993, Altan 5613 (GAZI). — C4 IÇEL: Abanoz-
Kazanci, 1300 m, 19.8.1994, Vural & al. 7154 (GAZI). — C6 GAZIANTEP: Antap [Gaziantep], Aucher-Aloy 1258 (P).
LEBANON: Antilebanon, bei Baalbeck, 16.10.1874, Dingler (B, M); in valle juxta Baalbak et ad Bescherre, 6.–7.1820-1826, 
Ehrenberg (LE); Barouk, 1.7.1937, Frér Louis (MSB, P); Ouadi el Harir, 16.7.1947, Mouterde (G); champs prés du Libani 
en face d’ Aytanit, 31.5.1955, Mouterde 11540 (G); Zachle, 1.7.1930, Gombault 1371 (P); Baalbek, 24.8.1930, Gombault 
1372 (P); Zachle, locus aridis, 25.8.1920, La Colte (L); Cedres de Barouk, 13.7.1937 and 18.7.1947, Mouterde 6144 and 
9036 (G).
SYRIA: Sehel, En Nebk, 4500', Davis 6457A (E); C. Zebdani in Antilibanon, c. 4000', 14.10.1874, Dingler (B); Mt Hermon, 
betw. Wadi Shiba and Wadi Jinnah, 1200-1500 m, 21.7.1924, Eig (MSB); Col du Baida, 2.7.1933, Frér Louis (P); Qatma, 
23.5.1934 and 25.5.1937, Frér Louis (P); Hamidieh, 19.4.1937, Frér Louis (MSB, P); Alep, Ansarieh, 10.5.1937, Frér Louis 
(P); Alep, 20.5.1937, Frér Louis (MSB, P) and 5.1936 (P); Azaz, 17.5.1937 and 25.5.1937, Frér Louis (P); Alep, Ain Zall, 
20.5.1937, Frér Louis (P); Hamma (Coelsyria), 17.7.1932, Gombault 1373 (P); Azaz, 25.5.1937, Gombault 4088 (P); Azaz, 
1500-2000', 20.-24.5.1955, Haradjian 4440 (G); collib. volcanices ad Alep, Michaux (P); Jabal Saouane, au N de Nebk, 
21.4.1954, Moutérde 11018 (G); Yabroud, 17.7.1952, Pabot (G); ibid., 20.10.1953, Pabot (G).
IRAQ: MRO: Erbil, Shaqlawa, 900 m, 8.5.1947, Gillett 8071 (US); in ditione pagi Penjwin, in apertis quercetorum, 1500 m, 
19.-20.6.1957, Rechinger 10526 (E, G, W); MSU: Nr. Sulaimaniya, on road to Qaradagh, 20.6.1959, Wheeler Haines 1506 
(E).
PALESTINE: Env. of Jerusalem, 31.5.1953, Orshan 14821 (MSB); Judean mts, Kefar Ezyon, 16.8.1935, Eig (E); Vasques de 
Salomon, IV.1889, Jouannet-Marie 495 (BR).
JORDAN: University campus, Al-Jubaiha, near Amman, 15.5.1973, Al-Eisawi 1491 (MSB); Transjordan, Ajlun, Wadi Ierkan, 
5.7.1941, Davis 3780 (E).
IRAN: PROV. W AZARBAIJAN: Khoi to Schahpur, Ghare Tapeh, 15.10.1948, Behboudi & Aellen 5577 (W). — PROV. E 
AZARBAIJAN: 47 km on the road from Mianeh to Khalkhal, before Zarjabad village, 1800 m, 20.6.1986, Maassoumi & 
Abouhamzeh 56880 (MSB). — PROV. LORESTAN: Dorud, 5500', 21.5.1940 and 6.6.1941, Koelz 15618 & 18090 (W); Bisheh, 
1200 m, 28.5.1937, Köie 715 (C); Bisheh, 50 km a Khorramabad orientem versus, c. 1200-1400 m, 14.-16.7.1948, K.H. & F. 
Rechinger 5689 (B, E, G, M, W); in monte Schuturunkuh, 25.6.1905, Strauss (B, LE). — PROV. MARKAZI: Arak, Shahzand, 
Sorouneh, Kuh-e Souroneh, 2000-2650 m, 17.7.1984, Mozaffarian 48286 (MSB); Sultanabad, ad Mowdere, 14.6.1905, 
Strauss (B, BM, E, LE); ibid., 19.6.1904, Strauss (W); in dit. urb. Sultanabad, m. Raswend, 9.1898, Strauss (B); Dit. 
Tefresch in montosis, 1898, Strauss (B, W). — PROV. BAKHTIARI: Dashtak to Cherry, 13.6.1973, Iranshahr & Moussavi 
15582 & 15589 (W).
NOT TRACED: Antilebano, incultis inter Zebedani et Rascheya, 24.6.1897, Bornmüller 559 (B); untere Region des 
Antilebanon, 4000-5000', 5.5.1914, Dingler (B); Machalé, 15.5. and 17.5.1937, Frér Louis (P); Serghaya, 26.6.1939, Frér 
Louis (P); Kurdagh, 5.1942, Frér Louis (MSB); [Lebanon] Beersheba, 1863-1864, B.T.Lowne (E); [Syria/Lebanon] Masret 
et Bach, 1800 m, 17.7.1952, Pabot (G); [Syria/Lebanon] 10 km Sednaya, 3.7.1953, Pabot (G); [Syria/Lebanon] Jebel 
Keniseh, 24.7.1900, Post (G); [Turkey] Bayazetskii sandzhak, sandzhak-Konisi-Ipi, 2.7.1916, Schischkin (LE); Pr. Djamal-
Edin, m. Dary-Dag, 11.6.1929, Schelkovnikov & Kara-Murza (LE). 



Astragalus sect. Pterophorus Bunge

1. Astragalus acicularis Bunge 

TURKEY: B5 KAYSERI: Ercyias Dagh, 7000', 18.6.1934, Balls 1416 (B); in montis Argaei reg. alpina, 2200-2500 m, 
18.6.1890, Bornmüller 2135, 2136 and 2136a (B); Develi to Kayseri, S Kayseri, 1780 m, 9.6.1992, Nydegger 46212 (MSB); 
Erciyas Dagi, Kuzuy Yamar, 2500-3000 m, 20.9.1993, Vural & al. 6979 (GAZI); ibid., 2300 m, 1.7.1988, Aytaç 2401 
(GAZI); 30 km to Devely from Kayseri, Erciyas Dagi, 2150 m, 9.8.1997, Zarre 39 (MSB); Erdschiadagh, auf steinigen 
Abhängen oberhalb Hadschilar, c. 2100-2500 m, 6.1902, Zederbauer (B, WU); Erdschiadagh, auf den Tcki-yaila, c. 2400 m, 
6.1902, Zederbauer (WU).


2. Astragalus baibutensis Bunge

TURKEY: A4 KASTAMONU: Kastamonu, Ilgaz Dagi, 5.9.1994, 2050 m, Adigüzel & al. 1783 (GAZI). — A4 ÇANKIRI: Isik Dag 
Zirvesi, volkanik anakaya üzerinde, c. 2000 m, 8.10.1975, Akmen 4004 (E). — A5 AMASYA: Wilajet Kastambuli 
[Kastamonu], Tossia, Kanak-Tschesme, 6.7.1892, Sintenis 4559 (B, BR, G, GOET, LE, M, MSB, P, WU); A8 GÜMÜSS#ANE: 
[Gümüs#sane] Stavri, 7500', 1.8.1933, Balls 537 (E); Gümüss#ane to Siran, 10 km E Siran, 1400 m, 18.7.1992, Nydegger 
46800 (MSB). — A8 RIZE: Rize - Bayburt, S Seite des Soganli Geçidi, 2300 m, 20.8.1987, Engel 155 (MSB); Rize, 15 km 
nach Ispir, 2050 m, 19.8.1987, Engel 152 (MSB). — A8 ERZURUM: Erzurum’un 60 km kbisi, Kuzgun Deresi günei, 2100 m, 
10.7.1976, Tatli 4726 (E). — A4 ANKARA: Gazi univ. Egrdim Fakultesi Bahcesi, 850 m, 25.5.1983, Vural 2226 (GAZI). — 
B4 ANKARA: Bestepe, 850 m, 3.6.1992, Vural 6018 (GAZI). — B6 SIVAS: 40 km N of Zara (71 km E Sivas), 1900 m, 
26.6.1969, Andersen & Petersen 16 (E, K). — B7 ERZINCAN: N Erzincan, Otlukbelidaglari, between Esekmeydani and 
Deverkorum, 2390 m, 17.7.1996, Rasmont 085 (LG).


3. Astragalus condensatus Ledeb.

TURKEY: A4 KASTAMONU: 5 km N Kastamonu, 900 m, 7.6.1954, Davis 21638 (E). — A5 ÇORUM: 15 miles from Çorum to 
Merzifon, 900 m, 30.5.1965, Coode & al. 1873 (E). — A5 AMASIA: Montis Ak-dagh, in regione alpine, 18.7.1889, 
Bornmüller 95 (B, BR: mixed with A. microptera). — B2 KÜTAHYA: Kesik Sögüt am Murat Dag, 1850 m, 4.6.1989, 
Nydegger 44066(MSB). — B3 KÜTAHYA: Sakivköy, c. 20 km SE Tavsanli, W Kütahya, 10.6.1989, Boratynski & al. 8126 
(MSB); Gediz, 850 m, Pinus bruttia forest, 4.7.1962, Davis & Coode 36892 (E, K); Gediz to Cavdarhisar, 1100 m, 4.7.1962, 
Davis & Coode 36999 (E, M); c. 3 km NE Kütahya along the main road, Cinarli Merkii, near Yazikanlar Petrol service 
station, c. 950 m, 21.5.1990, Lampinen 7608 (H). — C3 ESKISS#EHIR: Straße Mahmudyie - Gökcekaya, c. 500 m, S Mamure 
köyü an der Straße Alpu - Mahmudiye, 980 m, 2.6.1969, Buttler 13281 (M); N Eskisehir, 2.6.1955, H. & E. Walter 6 (B); 
Sündikendaglari, c. 1100 m, 20.6.1971, Ekim 1217 (E); Eskissehir - Sedigazi, in graminosis siccis prope Karabazar, 9.6.1932, 
Scheibe 1061 (B); Mahmudjeh - Ralkhugel, 20.4.1901, Warburg & Endlich 122 (B). — A4 ANKARA: Çubuk, Karagöl. P. 
nigra forest, 1600 m, 9.7.1974, Erik 636 (E); Ankara - Ayasdaglari, 1200 m, 6.6.1975, Akman 4007 (E); Ankara - Ayasbeli, 
19.6.1975, Akman 3098 (E); Polatli, Ankir area, 840-850 m, 6.6.1990, Aytaç & Duman 3072 (GAZI); Angora, champs 
abandonnés, Kodja-dagh, 4.6.1908, Aznavour 497 (G); ad Angora Galatiae, 1892, Bornmüller 3134 (B, W, WU); Galatia, in 
agro Ancyritano, ad montem Dikmen-dagh, 2.6.1929, J. & F. Bornmüller 13389 (B); steppes entre Nevessehir et Ankara, (82 
km avant Ankara), 1150 m, 15.7.1973, Cartier II2 (E); Ankara to Sirvihisar, 10 miles from Ankara, 850 m, 21.6.1962, 
Dudley 35995 (E); Kulu to Ankara, 10 miles from Kulu, open steppe hillside, 18.6.1962, Dudley 35977 (E); Steppe um 
Ankara, 20.6.1932, Kotte (M: mixed with A. plumosus subsp. plumosus); dry S slope in hills, Beynam, 55 km SW Ankara, 
12.7.1965, Ledingham & Ekim 4313 (E); dry overgrazed waste areas, W edge of Ankara, c. 850 m, 10.7.1965, Ledingham 
4294 (E) and 4298 (E, LE); SE Ankara, between Cakal Barla, 1200 m, 20.6.1968, Rix & al. 686 (E); Ankara, Garten der 
deutsche Botschaft, 29.5.1955, H. & E. Walter 3631 (B); Ankara, above Bala, 1400 m, 9.6.1967, Watson & al. 2614 (E). — 
C5 NIG#GDE: Çamardi - Demirkazik, 1580 m, 26.8.1993, Vural & al. 6863 (GAZI); 10 km östlich von Ulukisla (Vilajet 
Nigde), 12.6.1937, Reese (BASBG); Ala-Dag, Cemis Deresi, 2050 m, 21.8.1938, Ellenberg 101a (B); Ala Dag, NO Lager, 
2650 m, 25.8.1938, Ellenberg 1027 (B); Arpalik gorge NE Çamardi, 1700 m, 26.6.1989, Nydegger 44254 (MSB); Ala Dag, 
10.7.1964, Wood & Gibson 234 (E). — B5 NEVSEHIR: Kirsehir to Guls#ehir (Arapsun), c. 20 km from Gulssehir, c. 500 m, 
2.8.1956, McNeill 377 (E); Zelve, 1110 m, 22.5.1989, Vural & al. 4950 (GAZI). — B5 KAYSERI: Ali Dagh, Talas, 5000', 
17.6.1934, Balls 1382 (E); Barsania, 28.5.1868, Baytop 12767 (E); Bakir Dag between Kisge and Bakir Olugu Y., 1700 m, 
28.6.1952, Davis 19507 (E); Ali-Dag S Kayseri, E-Seite bei Talas, 1430 m, 10.6.1987, Nydegger 42716 (MSB); Kayseri to 
Kirss#ehir, 1 km to Muçur, 1120 m, 15.8.1997, Zarre 174 (MSB). — B6 KAYSERI: Sariz, Yalak (Yesilkent) Kuzeydogsu, 
Binbog#a Dagi, 1650-1800 m, 4.8.1988, Aytaç 2503 (GAZI); c. 40 km S Pinarbasi, on pass between Sariz and Carsak, 2000 
m, 22.6.1968, Rix & al. 662 (E). — B6 SIVAS: Pontus australis, ad Siwas, 1300 m, 6.5.1890, Bornmüller 4726 (B); Sivas-
Sarkisla, 1500 m, 28.8.1957, Davis & Hedge 32725 (E); Kayseri - Sivas, 11 km E Kayadibi, 1340 m, 13.6.1992, Nydegger 
46252 (MSB). — B7 SIVAS: 12 km N Zara, 1700 m, 12.7.1971, Andersen & al. 2188 (C); in aridis inter Caesaream et Siwas, 
1300 m, 12.6.1890, Bornmüller 2132 (B); Sivas - Sarkissla, 1500 m, 28.8.1957, Davis & Hedge D32725 (E); Kangal - Divrigi, 
5 km E Karasar Geçidi, 1550 m, 15.6.1992, Nydegger 46286 (MSB); Sarkisla - Agcakisla, 6.1954, Wohak (B); 12 km W 
Zara, 1300 m, 19.8.1971, Volk 71/592a (MSB). — C4 KONYA: Beyss#ehir to Konya, 30 miles from Beysehir, 1310 m, 
15.6.1962, Dudley 35866 (E); Sultanhani to Askil between Koglar and Halkendi, 1000 m, 17.6.1962, Dudley 35921 (E, M); 
Weidetrift bei Beysehir, 1000 m, 1.6.1974, Ern 140 (B, MSB); very dry hillside at Aegirmen Koy, 20 km W Konya, 
20.8.1970, Fraser Jenkins 2114 (E). — C4 KARAMAN: Karaman - Mut, 6 km S Karaman, 1080 m, 22.5.1993, Nydegger 
47260 (MSB); 19 km N Karaman, 14.6.1950, Reese (BASBG). — C5 IÇEL: 19 km N Arslanköy, 2000 m, 18.7.1970, 
Spitzenberger 90 (W).
NOT TRACED: Phrygia-Bithynia, Gebirge W Station Borüjuk, 7.1929, Borulard (B); Hadjin to Karakilisse, 9.7.1906, Post 267 
(G); Central Anatolia, Kargali, Camurcuk, 930 m, 6.6.1996, Lammont 579 and 731 (MSB); ibid., 1050 m, 10.7.1996, 
Lammont 791 (MSB); ibid., 6.6.1996, Lammont 583 (MSB); Lycia, inter Tschaipazar et Makri, 6000', 20.7.1883, Pichler 232 
(G); Asie mineure, 1855, Tchihatcheff (P).


4. Astragalus cymbibracteatus Hub.-Mor. & D.F. Chamb.

TURKEY: C5 NIG#DE: Nig#de-Çamardi, W Üçkapili, 1680 m, 24.7.1992, Nydegger 46834 (MSB). — B5 KAYSERI: Kayseri, 
N Saimbeyli, Obruk, 1520 m, 10.7.1995, Rasmont 089 (LG); c. 8 km to Tasçi from Devely, Bakir Dagi, beside the road, 1450 
m, 9.8.1997, Zarre 52 (MSB); 55 km to Saimbeyli from Tasçi, Bakir Dagi, 1500-1600 m, 9.8.1997, Zarre 56 & 57 (MSB). 
— B6 SIVAS: 42 km N Gürün, NW exposition, 1400 m, 1.7.1970, Sorger 70-33-54 (E). — B6 MALATYA: Malatya - Kayseri, 
40 km hinter Malatya, 1200 m, 25.8.1987, Engel 173 (MSB). — C5 IÇEL: SE Sariadin, 1440 m, 3.7.1995, Rassmount PR022 
(MSB). — B6 KAHRAMANMARAS#: Elbistan, Sar Dagi, 1500-1650 m, 9.8.1988, Aytaç 2550 (GAZI); Sariz, Binbog#a Dagi, 
Yalak area, 1500-2200 m, 5.8.1991, Aytaç & Duman 4417 (GAZI); Engizek Dagi, Biçmeoluk damlari çevresi, 1400-1500 m, 
23.7.1987, Duman 3686 (GAZI); Göksun - Elbistan, 6 km W Elbistan, 1090 m, 3.7.1989, Nydegger 44335 (MSB); 30 km to 
Göksun from Sariz, Binboga Dagi, 1700-1900 m, 10.8.1997, Zarre 74 (MSB); 25 km to Göksun from Sariz, Binboga Dagi 
area, 1600-1800 m, 10.8.1997, Zarre 102 (MSB); 29 km after Göksun to Elbistan, Çardak, Beryt Dagi, 2060 m, 12.8.1997, 
Zarre 127 (MSB).


5. Astragalus drusorum Boiss.

TURKEY: B6 KAYSERI: Cilicia, frequens inter Güllek et Boghas, 4800', 11.7.1853, Kotschy 99b (GOET, LE). — C2 IÇEL: 10 
km N of Ezerabat, 6.7.1971, Andersen & al. 2052A (E); env. of Gözne, NW of Mersin [Içel], 1000-1100 m, 16.8.1931, Eig & 
Zohary (M, MSB); Kiranh köyü-Madraneayt Kozak, 15.5.1966, Premeru 543 (B). — C6 GAZIANTEP: Roadside, hillside at 
Fevzypaza, 27.7.1971, Aberdeen Uni. Amanus Exp. Flora of Turkey (E); Nur Dagi Pass, W Fevzipasa, 1150 m, 24.6.1971, 
Sorger 71-37-11 (E). — C6 HATAY: Base du Mont Cassius, 5.7.1973, Cartier I (E); Okjilar village, between Antioch et El-
Ourdu, 4.7.1932, Eig & Zohary (E); Vallá de Karadourane (O. Kessab), 20.7.1955, Pabot (G).
LEBANON: Environs de Saida, 15.7.1853, Blanche (P); Bikfaya à Broummana, 26.6.1930, Blanche (Gombault) 1370 (MSB, 
P); in saxosis frutetorum ad Broummana, 700-800 m, 15.7.1897, Bornmüller 553 (B, BR, LE, M, MSB, P); in pinetis regionis 
inferioris ad Beit-Meri, 700-800 m, 20.6.1897, Bornmüller 554 (B, BR, E, G, LE, P) and 17.7.1897, Bornmüller 555 (B,); in 
pinetis regionis inferioris ad Brummana, 700-800 m, 21.7.1897, Bornmüller 556 (B, BR, E, G, LE, P); Montium Libani 
australis in declivitatibus occ., in pinetis ad Ain Zahalta, 1200-1300 m, 17.6.1910, J. & F. Bornmüller 11584 (B, E, G, LE, 
W); Montium Libani australis in dec. occ. ad pagum Bhamdun, bei Drusendorf, 5.6.1910, 1200-1300 m, J. & F. Bornmüller 
11585 (B); in Felsen bei Rhazir, c. 400 m, 18.6.1934, Busujan 153 (M); juste avant Meshta Helon, 500 m, 7.7.1973, Cartier 
II (E); Bairut, 5.1820-1826, Ehrenberg (BR, L, LE, P); Deir el Kamar, 29.6.1973, Frér Louis (P); Keyfour, 1.7.1938, Frér 
Louis 332 (MSB, P); champs en jachére prés de Berté [Barti], á l'est de Saida, 25.7.1858, Gaillardot 1033 (B, LE); Ain 
Zehalta, 10.8.1933, Mouterde 2752 (G); Dekoané, 14.5.1936, Mouterde 5230 (G); Qortada, 2.7.1950, Mouterde 9898 (G); 
Süg-el Gharb, 5.1900, Post (G).
SYRIA: Djebel Druse, 15.7.1920, La Cotte (L); ibid., 20.10.1953, Pabot (G); entre Antioch et Souriyé, 14.8.1933, Thiébaut 
T18 (G).


6. Astragalus kirshehiricus D.F.Chamb.

TURKEY: B4 KIRSEHIR: At the entrance of the city from Ankara, low hills near the fire brigade, 1040 m, 8.8.1997, Zarre 21 
(MSB). — B4 AKSARAY: Kizilkaya köyü, Karaya merkii, c. 1100 m, 21.9.1994, Ertug 223 (GAZI). — C4 KONYA: 5 km S 
Karapinar, 12.7.1971, Andersen & al. 2166 (E); 3 km from Cihanbeyli to Tuz Gölü, 900 m, 18.6.1962, Dudley 35973 (E); 
entrance to Topraksu Govt station near Karapinar, c. 100 km E Konya, dry sandy wind eroded plain, c. 1000 m, 16.7.1965, 
Ledingham & al. 4367 (E).


7. Astragalus lesbiacus Candargy

GREECE: EAST AEGEAN ISLANDS: CHIOS: N slopes of Pelinaion, open sunny places, 400-800 m, 14.5.1985, Boratynski & al. 
54 (E); loose Pinus brutia forest between Ag. Geogrios and the cross-road toward Anavatos and Lithi, 31.5.2985, Boratynski 
& al. 211 (E); N slope of Orors Mt climb from Kardamila, open stony places, c. 700 m, 28.5.1985, Boratynski & al. 184 (E); 
s. loc., 1908, Fréres des E. C. 50 in Hb. Aznavour (G); 2 km w Agios Isidoros, c. 400-500 m, 13.6.1966, Lüdtke 560 (M); s. 
loc., Olivier & Bruguiére (P); Chios, 1.5.1902, Sterneck 176 (MSB, PRC). — LESVOS: The eastern hill by Karionas village, 
W Skopelos, 800 m, 23.5.1993, Hansen & Nielsen 8737 (C). — SAMOS: 2 km N summit prov. Ilias of Mt Ampelos, near the 
road from Mon. Bronti, 800-900 m, 23.6.1980, Baden 449 (C); s. loc., 15.6.1886, Forsyth Major 468 (G); about half way 
between Arvanitis and Vourliotis under scattered pine trees, 3000', 25.6.1963, Gathorne-Hardy 537 (E).
TURKEY: B1 MANISA: In mt. Sypilo, 1837, Aucher-Eloy 1260 (G, P); partie supérieure du Sipyle, au dessus de Magnésie, 
17.6.1854, Balansa 214 (E, G, GOET, LE, MSB, P, W); in mte. Sipylos, supra Magnesia, 800-900 m, 10.6.1906, Bornmüller 
9413 (B); Manisa Dagi, Islik Deresi, 1200 m, 19.6.1984, Duman 1779 (GAZI); Sipylos Magnesiae, in lapidosis summis 
montis, c. 1600-1800 m, 12.8.1933, Schwarz 1003a (B); Manisa Dagi (mt. Sipylus), 1450 m, 5.7.1984, Strid & al. 23234 (B, 
C, G). — B1 BALIKESIR: Kaz Dag, 1600 m, 27.6.1977, MacPhail & Watson (E). — B1 IZMIR: Kempalpasa, in forest duff in 
Pinus bruttia zone on the lower slopes of Nif Dag, 30.5.1966, Alava & al. 4880 (E); Sabounjou-Kaive, entre Smyrne et 
Magnésie, dans les terrains calcaires, 9.6.1854, Balansa 216 (G); Nifdag, in pinetis apertis, c. 600-1000 m, 12.5.1933, 
Schwarz 650 (B); ibid., c. 1000-1300 m, 7.7.1933, Schwarz 815 (B). — C3 ANTALYA: Tahtali Dag (c. 50 km SW Antalya), 
W side (ascent from the village of Yukari Beycik), 2100 m, limestone, 17.7.1984, Strid & al. 23623 (B, G); Bunarbaschi, in 
mte. Takhtali dagh, 500-700 m, 26.5.1906, Bornmüller 9412 (B, G, P, W); ibid., 6.1932, Schwarz 336 (B).
NOT TRACED: Asia Minore, Mykali, 25.6.1886, Forsyth Major 716 (G); mountain side above Gölü, 14.7.1940, Post (G).


8. Astragalus parnassi Boiss.
8a. Astragalus parnassi subsp. parnassi

“MACEDONIA”: Gradsko, ad fluvium Warder, 200 m, 22.5.1917, Bornmüller 763a (PRC); Wardartal bei Stobi, mittl. 
Mergelterasse, 150 m, 30.5.1971, Breckle 1470 (M); Steppe bei Stip, 5.1918, Burgeff (M); Steppe bei Krivolak, 22.5.1918, 
Burgeff 426, 226 (M); an der Autostr. Thessaloniki - Nis. Kalkmergelhänge hinter der Abzweigung nach Kavardaci, c. 80 km 
N der Grenze, 28.6.1985, Erben (MSB); Titov Veles, 5.-6.1972, Klaus & al. (H); in collinis steposis ad Sv. Nikula (Ovce 
Polye), 19.5.1926, Košamin & C#Cernjavsky (LE); an der Autostr. Thessaloniki - Nis, Kalkmergelhänge hinter der Tankstelle 
an der Abzweigung nach Kawardazi, c. 80 km N der Grenze, 28.6.1985, Lippert 21047 (M, MSB); inter Gradsko et 
Chichevo, in locis lapidosis et graminosis, c. 250 m, 18.5.1963, Mayer 50592 (M); Topolka-Kloof, S Titov Veles, 23.5.1963, 
Ooststroom 23262 (L); N-S Streichende Ausläufer des Plevenski Rid W Titov Veles, 400-600 m, 4.8.1976, Podlech (MSB).
ALBANIA: M. Murga, prope frutem flamburas supra Puli, distr. Paramythia, 6.7.1895, Baldacci 35 (LE, P, PRC).
GREECE: ATHENA: pr. Athenas, in collibus saxosis, IV.1848, Heldreich (P). — THESSALIA: Ossa, N Seite, alpine Matten, c. 
1900 m, 23.6.1981, Erben (M, MSB); M. Ossa, 7.1884, Heldreich (W); 6 km NE Spilja, 940 m, 4.6.1972, Klaus & al. 193 
(W); Larissa, S-Hänge des Ossa bei Anatoli, 1200 m, 18.6.1974, Raus 2269 (MSB). — VIOTIA: In Parnassos, inter Zeitra et 
Solano, Berger (M); Westabfall des Parnassos in der oberen Abies cephalonica-Stufe, 26.5.1969, Bresinsky (M); Parnassos, 
steinige Wiese an der Straße zur Schutzhütte, c. 1300 m, 18.6.1981, Erben (M, MSB); Parnassos, in der Umgebung des 
Kataphygion, c. 1550 m, 18.6.1981, Erben (herb. Erben); Parnassos, 1840, Fraas (M); m. Parnes, 5.1930, Gioul (BC, M); S-
Hänge des Parnassos, östlich Arachow an der Straße Livadia-Delphi, 17.5.1967, Grau (MSB); in m. Parnassi, 7.1855, 
Guicciardi (P); in oropedeia Keramitia m. Parnethis supra Khasia, 3000', 5.6.1857, Heldreich 615 (LE, M, P, WU) and 
22.6.1896 (LE, P); in m. Parnethe prope Dekleiam, 15.6.1878, Heldreich (P); in m. Parnethis reg. media, 18.6.1898, 
Heldreich (E, M, W); Straße Livadia - Delphi, S-Hänge des Parnassos, östlich Arachow, 900 m, 14.-18.5.1967, Kress (M); 
ibid., Haesler 1688 (LE); Parnass, Tannenregion, 1500 m, Lemperg 629 (W); in m. Parnethis reg. media, 18.6.1898, 
Heldreich (E, M, W). — PHOKIS: Above Ypati, Oeta, 4500', 9.6.1937, Balls & Balfour Gourlay 3190 (E); Hänge bei Delphi, 
22.4.1954, Merxmüller & Wiedmann 10745 (M).
NOT TRACED: M. Epanokrepa, 2000', 1834-1842, Fraas (M).


8b. Astragalus parnassi subsp. calabricus (Fisch.) Zarre

ITALY: SICILIA, CALABRIA, PROV. DI COSENZA: S. Giovanni, 1200 m, 25.7.1910, Lopez (WU); Silla en Calabra, 1816, coll. 
ignatus (P); Sila Grande, bei Spezzano di Sila, c. 1000 m, 14.8.1965, Merxmüller & Grau 20609 (M).


9. Astragalus phrygius Širj.

TURKEY: C3 ISPARTA: Dedegöldaglari, Kizil Dag National Park, open stony place, 1300-1400 m, 27.5.1991, Boratynski & al. 
6869 (MSB); Sultandaglari, an der Straße von Akssehir Richtung Gelendost, knapp E unterhalb der Paßhöhe zwischen Engilei 
und Örkenez (= Bagkonak), 1600-1650 m, 10.7.1978, Ehrendorfer & al. 787-21-4 (WU); Paßhöhe SW Akssehir in den 
Sultandaglari, 1600 m, 7.7.1973, Holtz & al. 549 (E); Isparta-Akssehir, in Paßhöhe Sultandaglari, 1710 m, 15.6.1989, 
Nydegger 44176 (MSB).


10. Astragalus strictispinus Boiss.

TURKEY: C2 MUG#LA: Çal Dag (N Fethie), S side, 1800 m, rocky slope in opening of mixed coniferous forest (Pinus nigra, 
Cedrus libani), 9.7.1984, Strid & al. 23395 (G); Köycegiz, Agla-Eskere arasi, 1700-1900 m, 25.7.1992, Güner & al. 10750 
(GAZI). — C3 BURDUR: Waste ground along the road, 15 km S Dinar on the way to Burdur, 15.6.1966, Alava & Bocquet 
5197 (E); Pisidia, Vilayet Burdur, 10.6.1938, Reese (BASBG); Vilajet Afyonkarahisar, 14 km S Dinar, 12.6.1938, Reese 
(BASBG). — C2 ANTALYA: Distr. Elmali, Bey Dag, 2000 m, 28.7.1960, Khan & al. 304 (E); auf dürren Hügeln W 
Abhängen bei Elmali, 20.7.1883, Pichler (WU); Ciglikara forest S of Elmali, open places in a cedar forest, 20.6.1989, 
Boratynski & al. 8120 (MSB).


11. Astragalus talasseus Boiss. & Balansa

TURKEY: B5 NEVESSEHIR: Göreme-Avanos, 940 m, 20.6.1989, Vural 5306 and 5554 (GAZI); Cavusin - Avanos, 1000 m, 
9.8.1989, Vural & Eyüboglu 5548 and 5554 (GAZI, MSB). — B5 KAYSERI: mit lückiger, niedriger Vegetation bestandene 
Hügel am Ortsrand von Ürgüp, 1100 m, 4.10.1989, Hertel 35942 (M); in collibus supra Cesareae, 23.9.1904, Maire 612 (B); 
Yilanli Dagi, 1150-1250 m, 22.9.1993, Vural 7041 (GAZI); in arenosis ad litus occidentale stagni salsi S Yeshilhisar, 1100 
m, 26.7.1974, Rechinger 49956 (M, W); c. 8 km to Tasçi from Devely, Bakir Dagi area, near the road, 1450 m, 9.8.1997, 
Zarre 51 (E, MSB).


12. Astragalus thracicus Griseb.
12a. Astragalus thracicus subsp. thracicus

GREECE: MAKEDONIA: 1837, Aucher-Eloy (P); environs de Salonique, 15.8.1917, Charier (P); pag. Kereci-Köj pr. 
Thessalonicam, 9.1904, Dimonie (M); circa Thessalonicam, 1820, Dumont d'Urville (P); Chalkidike, Halbinsel Kassandra, an 
der Küste bei Athine, N Kallithea, 9.7.1984, Erben (M, MSB); Halbinsel Kassandra, an der Küste bei Athine N Kallithea, 
10.7.1984, Erben & al. (herb. Erben); inter Galacinto Chereridire et Salonicki, 6.8.1871, Janka 44 (GOET); in campis aridis 
ad vias pr. Thessalonicam versus m. Karthiati, 10.7.1851, Heldreich 2340 (W); pr. Thessalonicam, 2.-14.7.1857, Orphanides 
599 (LE, M, P); in cacumine montis Malevo Laconiae, 6700', 7.-19.6.1857, Orphanides 600 (BR); Kaputsidees prope 
Thessaloniki, in collibus siccis, 5.6.1936, K. H. & F. Rechinger 9006 (M, W); Saloniki, 3.10.1918, coll. ignatus (P). — 
THRAKIA: 2-3 km W Pherai, in Quercus coccifera scrub, 20.6.1979, Boratynski & al. 487 (MSB).
TURKEY: A1 TEKIRDAG: Telki Dagh [Tekirdag], in collinis secus litora Oropontides inter Rodasto et pagum Kumbaos, 
20.6.1890, Degen (WU). — A2 ISTANBUL: Küçükçekmece to Büyükcekmece, 50-100 m, 12.8.1962, Davis & Coode 39274 
(E). — B1 IZMIR: Yamanlar Dag, Karagöl, Pinus brutia forest, on the hillside above the lake, 9.6.1966, Alava 5056 (E); 
Smyrne, Aucher–Eloy 1252 (MSB); Sabounjou - Kaive, entre Smyrne et Magnésie, dans les terrains calcaires, 19.6.-
18.8.1854, Balansa 216 (BR, G, GOET, MSB, P); in Troade, Bruner (M). — B2 USS#AK: Phrygia, Ouchak (phrygie), vers 
910 m, 6.1857, Balansa (P). — B3 BILECIK: Pazaryeri, 750 m, Steppe, 2.7.1962, Davis & Coode 36500B (E); Abaslyk, bei 
Biledschik, 26.7.1873, Dingler 98 (B). — B3 ESKISS#EHIR: 35 km on the road Eskissehir - Porsuk, Bozuk, Quercus forest, 900 
m, 23.7.1993, Aytaç & Ekici 6204 (GAZI); Karaköi, 8.1906, Schinabeck (B, M).
NOT TRACED: 10 km N Ezerabat, 6.7.1971, Andersen & al. 2052 (E).


12b. Astragalus thracicus subsp. monachorum (Širj.) Strid

GREECE: MAKEDONIA: M. Athos, 1867, Middendorf (LE).


12c. Astragalus thracicus subsp. afyonicus Ponert

TURKEY: A4 ANKARA: Ad versuras 48 km NW Sereflikochisar, 27.7.1974, Rechinger 49978 (MSB, W).


13. Astragalus tmoleus Boiss.

TURKEY: B1 IZMIR: 10-11 km NE Ödemis, on 7th km of the road from Birgi to Salihli, 2.7.1989, Boratynski & al. 8123 
(MSB); Boz Dag, between Abbas and Geylan yayla, 1650 m, 16.7.1947, Davis 13318 (E, M); Yamanlar Dag, 27.4.1969, Fitz 
& Spitzenberger Sp-266 (W). — A3 BILECIK: Bei Biledschik, 10.7.1873, Dingler 99 (B). — B3 ESKISS#EHIR: 31 km to Porsuk 
from Eskiss#ehir, 1000 m, 23.7.1993, Aytaç & Ekici 6200 (GAZI); Eskisehir, Tasohek deposi, c. 1200 m, 1.7.1970, Ekim 593 
(E); Sündiken Dagi, Atalan tekke, 1000 m, 21.6.1974, Ekim 903 (E). — A3 ANKARA: Beypazarinin batis, Korunmus marnli 
saha, 800 m, 21.7.1993, Aytaç & Ekici 6127 and 6130 (GAZI). — C2 DENIZLI (CARIA): Babadag, Pinus bruttia forest, 1200 
m, 29.7.1988, Aytaç 2454 (GAZI); en montant au Babadag (au dessus de Denizli), 1500 m, 17.7.1973, Cartier 2 (E); Elmali, 
23.7.1883, Pichler 14 (G). — C2 BURDUR: 10 km to Dirmil from Gölhisar, 1000-1200 m, 23.8.1993, Duman & 
Karaveliogullu 5298 (GAZI); Esenli köy, Sertac-Burdur, 22.7.1968, Gözler 14161 (E). — C3 ISPARTA: Gelincik köy-Isparta, 
7.8.1968, Gözler 14161 (E). — C3 ANTALYA: Gömbe köy, Kas, 6.8.1968, Gözler 14151 (E); Beuge bei Karaköi an der 
Anatolien-Bahn, 8.1906, Schinabeck (B, M); Calbali Dag, SE part of Bakirli Dag, 1550-1650 m, 18.7.1984, Strid & al. 23677 
(MSB). — C4 KONYA: Ermenek, Tekeçati arasi, Açik yerler, c. 1450 m, 7.7.1978, Vural 965 (GAZI). — C6 ADANA: 
Hasanbeyli, Almanpinari, c. 1000 m, 21.7.1988, Aytaç 2450 (GAZI). — C4 IÇEL: Mut, Alahan, c. 700 m, 18.6.1979, Vural 
1516 (GAZI); Mut, Palantepe, 240 m, 25.8.1992, Vural & al. 6076 (GAZI).


14. Astragalus verae Širj.

TURKEY: A3 BOLU: In monte ad orientem lacus Abant, 1600-1750 m, 28.7.1974, Renz 50060 (MSB, W); Köroglu Dag, 
Felsplatten und alpine Festuca-Rasen auf einem Nebengipfel SÖ des Hauptgipfels, 2100 m, 11.8.1982, Raus 7016 (B); 
mountain above Mundurnu, bare banks, 9.7.1962, Davis & Coode 37104A (E); Hänge am Abant Gölü, SW Bolu, c. 1350 m, 
25.9.1957, Wagenitz & Beug 235 (B).


15. Astragalus yuralicus Boiss.

TURKEY: B5 KAYSERI: Zelve (Höhlenstadt bei Göreme), N Ürgüp, 21.8.1969, Albertshofer (M).