Instructions for authors

Willdenowia — Instructions for authors (August 2018, adjusted April 2019)

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Willdenowia is an international peer-reviewed journal publishing original research papers in English from the entire fields of plant, algal and fungal systematics, covering the evolution, taxonomy and nomenclature of these organisms as well as related fields such as floristics and plant geography. Papers on phylogeny and molecular systematics are especially welcome, as are review papers. Descriptions of new taxa are considered when supported by robust evidence. Floristic notes are generally not considered, except as contributions to the Euro+Med-Checklist Notulae series. Authors are encouraged to deposit duplicates of their material, especially nomenclatural types, at the Berlin herbarium (B).

Willdenowia is published in an open-access online edition and a print edition. Authors of articles retain copyright under the Creative Commons Attribution 4.0 International licence (CC BY 4.0), which permits unrestricted use provided the original author and source are credited (see

Manuscripts submitted for publication must be accompanied by a written and signed statement confirming that (1) neither the manuscript nor parts of it have been published before or submitted for publication elsewhere, and (2) it has been read and approved by all authors. Material, e.g. illustrations, already published elsewhere is not normally accepted for publication; exceptions are subject to any required permission from the copyright holder already having been obtained by the author.

Manuscripts must conform to the style and editorial standards of Willdenowia. Manuscripts are submitted by e-mail as a single DOC, DOCX or RTF file. Illustrations are initially accepted as low-resolution images and may be embedded in the manuscript or sent as separate files.

Manuscripts are subject to peer review by experts in the field. The author for correspondence receives the decision on acceptance or otherwise of a manuscript from the Editor immediately after the conclusion of the peer-review process. Authors may be asked to revise their manuscripts to accommodate recommendations of the reviewers and the Editor.

When preparing a manuscript, consult the most recent issues of Willdenowia (either the print edition or the PDF files in the online edition – not the HTML text) and follow this format strictly. British (U.K.) English spelling is used. In particular, be sure to conform with the following standards:

The manuscript must be headed by a concise and informative title and the full name(s) of the author(s) and their address(es); if there is more than one author, identify the author for correspondence. An abstract of up to 200 words followed by up to 15 key words precede the main text. The main text should be followed by acknowledgements, references and captions for the figures and tables.

Scientific names of taxa must be in accordance with Turland N. J. & al.: International Code of Nomenclature for algae, fungi, and plants (Shenzhen Code), 2018 (online at and are, irrespective of rank, consistently printed in italic type, as recommended by the Code. Personal names are not written in (small) capitals. Authors of scientific names (except above the rank of genus) are given at the first occurrence of such a name and are abbreviated in conformity with Brummitt R. K. & Powell C. E.: Authors of plant names, 1992, and its updates online (, except that spaces are used after full stops (periods), e.g. R. Br., W. D. J. Koch, L. f., Müll. Arg.

Synonymies and type citations appear in a standardized format; please consult recent issues of Willdenowia (see, in particular for the arrangement of heterotypic (taxonomic) and homotypic (nomenclatural) synonyms and their types.

Titles of serials in bibliographic citations following scientific names are separated from the author citation by “in” and abbreviated in conformity with Bridson G. D. R. & al.: BPH-2. Periodicals with botanical content. Constituting a second edition of Botanico-Periodicum-Huntianum 1–2, 2004 (online at Titles of serials not listed in BPH-2 are abbreviated according to the abbreviation standards there defined.

Titles of monographs are separated from the author citation by a comma and are abbreviated in conformity with Stafleu F. A. & Cowan R. S.: Taxonomic literature, ed. 2, 1–8, 1976–1988; & successors, suppl. 1–8, 1992–2009 (online at, except that all components start with capital letters. Titles of monographs not listed in TL-2 follow the version used in IPNI (

Label data of herbarium specimens, when standardized rather than cited literally, are given in the following sequence: Country, political subdivision or region, locality, , date, collector(s) collection number (herbarium code barcode/accession number). Standard herbarium codes follow Thiers B. 2018+: Index herbariorum ( For example: China, Yunnan, Gongshan Xian, about 4 direct km S of Gawagapu mountain and c. 15.9 direct km WSW of Bingzhongluo, 27°58'29"N, 98°28'29"E, 3880 m, 25 Aug 2006, Gaoligong Shan Biodiversity Survey 31398 (CAS 1089187).

When permits are required for collecting and exporting samples and conducting research, these should be cited in the Acknowledgements or under Material and methods.

Figures (Fig.), including drawings, photographs, diagrams and maps, are numbered with a single run of Arabic numerals in each paper. The same applies to tables (Table). Scale must be indicated in illustrations; please draw in scale bars and give the corresponding measurements in the caption. Elements within figures are lettered A, B, C, etc.

Identification keys conform to the following example (other formats, e.g. indented keys, are not accepted):

1.  Petal limb with a dark spot at base; capsule membranous; basal leaves fleshy or succulent, subglabrous ......................................................................................................................  1. S. sedoides
–   Petal limb without a dark spot at base; capsule relatively firm and opaque, rarely submembranous; basal leaves fleshy, sparsely hairy ......................................................................................................................  2
2.  Capsule 6 mm long or more, oblong; cauline leaves obovate to ovate; anthers lilac; hyaline margin of calyx teeth smooth or shortly and sparsely hairy ...................................................................  6. S. integripetala
–   Capsule 6 mm long or less, ovoid or oblong; cauline leaves spatulate to obovate; anthers white or pale lilac; hyaline margin of calyx teeth ciliate-fringed and often glandular .................................................  3

Bibliographical references in the text are cited in the following form: Webb & Camarasa (1972) or (Webb & Camarasa 1972); if the page number is stated, Webb & Camarasa (1972: 267); or, if there are more than two authors, Filipowicz & al. (2012). All bibliographical references quoted in the text, and only those quoted, are listed at the end of the manuscript, under the heading References, in a format strictly analogous to that of the examples below. This also applies to references quoted in an abbreviated way in the synonymy of taxa; but in papers with very many such references, this should apply only to recent references and to those citing numerous names, in order to avoid an unacceptable expansion of the references section.

References at the end of the manuscript are cited according to the following examples. For further examples see recent articles in Willdenowia (either the print edition or the PDF files in the online edition – not the HTML text). References are cited in the original language and, in case of non-Latin script, either in that script or transliterated; additional translations, if appropriate, are enclosed in square brackets. Titles of serials are abbreviated in conformity with BPH-2 (see above); titles of monographs are cited in full.

Filipowicz N., Nee M. H. & Renner S. S. 2012: Description and molecular diagnosis of a new species of Brunfelsia (Solanaceae) from the Bolivian and Argentinean Andes. – PhytoKeys 10: 83–94.
Kükenthal G. 1909: Cyperaceae – Caricoideae. – In: Engler A. (ed.), Das Pflanzenreich. Regni vegetabilis conspectus IV. 20 (Heft 38). – Leipzig: Wilhelm Engelmann.
Linnaeus C. 1753: Species plantarum. – Holmiae: Laurentii Salvii.
Thiers B. 2018+ : Index herbariorum: a global directory of public herbaria and associated staff. New York Botanical Garden’s virtual herbarium. – Published at .
Webb D. A. & Camarasa J. M. 1972: Odontites Ludw. – Pp. 266–269 in: Tutin T. G., Heywood V. H., Burges N. A., Moore D. M., Valentine D. H., Walters S. M. & Webb D. A. (ed.), Flora europaea 3. – Cambridge: University Press.

After acceptance of the manuscript, submit illustrations either as originals, mounted on individual sheets, or as graphic files (in TIF format, preferably never previously saved in JPG format). Graphic files may be (1) greyscale or colour illustrations with a resolution of at least 400 pixels per inch, or (2) line illustrations in bitmap mode (i.e. black and white only, not greyscale or colour) with a resolution of 1200 pixels per inch, in each case using a size at least equal to the size to be printed. Illustrations and tables including the space needed for the caption must, after reduction, fit into the available area on a page (16.5 × 24 cm). Colour illustrations are reproduced in colour in the online edition, and in either colour or greyscale at the Editor’s discretion in the print edition.

A single set of proofs is sent once and, except upon contrary instructions, to the author for correspondence. The author is expected to check and correct them carefully and return them without delay.

Printed reprints are not produced. Instead, the author for correspondence receives the DOI (Digital Object Identifier) of the open-access online article, where a PDF file may be downloaded. The authors (who retain the copyright under the CC BY 4.0 licence) may make the paper freely available by self-archiving, individually, institutionally, or in a subject-based open-access repository.

Correspondence to:

The Editor of Willdenowia
Nicholas Turland
Botanischer Garten und Botanisches Museum Berlin
Freie Universität Berlin
Königin-Luise-Str. 6-8
14195 Berlin

Tel.: +49 (0)30 838 50137