ISIDRO MÉNDEZ
The problem of Citharexylum spinosum (Verbenaceae) in the Antilles
Electronic supplement to:
Méndez, I.: The problem of Citharexylum spinosum (Verbenaceae) in the Antilles. – Willdenowia 31: 419-424. 2001. – ISSN 0511-9618.
Specimens examined
Citharexylum spinosum L. f. spinosum
Cuba: Provincia de Pinar del Río: San Diego de Los Baños, 31.8.1910, Britton, Earle & Gager 6864 (US); Sierra Mendoza, 25.12.1911, Shafer 11139 (US); vicinity of Sumidero (in dry rock), 28.7.1912, Shafer 13449 (US); Sierra Mendoza, 25.12.1911, Shafer 11139 (US). — Provincia de La Habana: Sierra de Anafe, 18.12.1911, Wilson 11402 (US); at río Almendárez, Habana, 10.11.1921, Ekman 13473 (S). — Provincia de Matanzas: Boca del Canasí, 18.12.156, Morton 10224 (US). — Province of Cienfuegos: Soledad, Cienfuegos, 29.6.1929, Jack 7359 (C, F); ibid., 29.6.1929, Jack 7359 (S); ibid., 2.4.1926, Jack 4538 (S). — Provincia de Santa Clara: 28.9.1927, Jack 5514 (S). — Provincia de Camagüey: Cayo Romano, camino de El Molino a Sabanita del Burro, 17.5.1976, Areces & al. 31671 (B); Pueblo de Romano, Cayo Romano, 8.10.1909, Shafer 2471 (F); elevaciones frente a la Playa del Muerto, ladera noreste de la Silla de Romano, Cayo Romano, (matorral costero), 18.2.1990, Risco, Caballero & Parrado 6904 (HIPC); vicinity of Romano, Romano Cay, 8.10.1909, Shafer 2471 (US); trocha a continuación del molino de viento, Cayo Romano (bosque siempreverde micrófilo sobre caliza), 17.2.1990, Risco & al. 6900 (HIPC); camino hacia el campamento Camagüey, Cayo Guajaba (manigua costera), 12.2.90, Risco & al. 6811 (HIPC); Cayo Ballenato del Medio, Bahía de Nuevitas (vegetación costera sobre carso), 4.8.1990, Morales, Plaza & Primellez 5582 (HIPC); costa norte de Cayo Sabinal (manigua costera sobre carso), 8.1984, Méndez & Trujillo 958 (HIPC); Playa de Santa Lucía, 19.3.1909, Shafer 955 (F); Paso de los Paredones, Sierra de Cubitas (vegetación de la cima de elevaciones calcáreas), 28.1.1988, Caballero & Risco 3793 (HIPC); camino entre Triffin y carretera a Cayo Sabinal (21º34'40''N y 77º20'15''W, bosque siempreverde micrófilo degradado), 3.3.2000, Greuter & al. 25164 (B, HAJB, HIPC). — Provincia de Las Tunas: Las Nuevas (bosque sobre redzinas rojas), 4.8.1990, Verdecia 8014 (HIPC); carretera de Manatí a Puerto Manatí, cerca de Los Pitos (21°21'40''N y 76°57'10''W, matorral xeromorfo espinoso alto sobre serpentina), 5.3.2000, Greuter & al. 25280 (B, HIPC, HAJB); Playa de La Boca, Puerto Padre (in manigua costera), 20.5.1957, López Figueiras 2952 (US); Playa Herradura, costa norte, manigua costera, 14.1.1988, Méndez & Verdecia 4015 (HIPC). — Provincia Holguín: Los Cocos de Gibara, camino de Gibara a Caletones, km 2 (matorral xeromorfo costero y subcostero sobre carso), 27.3.1990, Méndez & Parrado 6083 (HIPC). — Provincia de Santiago de Cuba: Juraguá, 8.1929, Bucher 1a (F); 24.1.1902, Pollard & Palmer 28 (F); Loma Vigía, Nagua, Sierra Maestra (in forest), 4.8.1922, Ekman 14727 (S); Paso Estancia, Arroyo Blanco, ad flum. Cauto (in sylva), 10.11.1921, Ekman 6268 (S). — Provincia de Guantánamo: Baracoa, Abra del río Yumurí (sobre roca caliza), 11.8.1975, Álvarez & al. 27324 (B); Baracoa, 24.1.1902, Pollard, Palmer & Palmer 28 (S); South Montecristo (in coastal limestone bluff), 16.1.1956, Morton & Alain 9194 (S); vicinity of Baracoa 18.2.1910, Shafer 3934 (US); Río Yumuri, 6.12.1910, Shafer 7167 (US); Wright 438 (S, US); Cueva del Agua, primer nivel de terrazas al noreste de Punta de Maisí (bosque siempreverde sobre carso), 4.8.1992, Méndez, Barbadillo & Ramírez 7077 (HIPC); Jobo Claro, Carretera de Baracoa a Maisí (matorrales secundarios sobre caliza), 3.8.1992, Méndez, Barbadillo & Ramírez 7277 (HIPC).
Jamaica: St Thomas Parish, 14 miles SE of Kingston toward Morant Bay, 6.1.1972, Wunderlin 5137 (F); Long Mountain (100 ft), 9.11.1904, Harris 8840 (F); Clarendon, north slope of Round Hill (dry limestone woodland), 500-1000 ft, 22.11.1954, Proctor 9506 (F); Port Henderson, Parish St Catherine, 5.2.1961, Adams 8884 (M); near Spur Tree, W. Don Figueredo Range, (c. 2500 ft, on rocks slopes), 23.1.1958, Yunker 18054 (S); 5 miles W of Port Santa María (c. 50-100 ft, in thicket), 26.12.1957, Yunker 17842 (S); near Port Antonio, 17.7.1897, Fredhoh 3169 (US); Great Goat, Old Harbour Bay, 4.3.1908, Harris 10169 (US); N slopes of Round Hill (500-1000 ft dry limestone woodland), 22.11.1954, Proctor 9506 (US); Clarendron Parish, road from Spaldings to Thomson Town (900 m, forest in limestone formation), 24.7.1997, Acevedo & al. 9659 (US); Trelawny, one mile of the coast at Falmouth Low (dry rocky land), 23.7.1952, West & Arnold 783 (C); Parish Manchester, second growth along roadside about 2 miles west of Spur Tree (about 2000 ft), 23.8.1965, Hespenheide 1519 (F); Long Mountain (700 ft), 9.11.1904, Harris 8840 (F); Old Harbour Bay, Great Goat, 4.3.1908, Harris 10169 (F).
Bahamas: Great Inagua, just northwest of Morton Bahamas headquarter (in open coppice), 23.6.1978, Correll 49939 (F); Inagua, 12.10.1904, Nash & Taylor 935 (F); Caicos Islands, N Caicos, Kew and vicinity, 18.12.1907, Wilson 7730 (F); Mariguana, Abraham Bay and vicinity, 6.12.1907, Wilson 7517 (F); E Caicos, Jacksonville and vicinity (in rocky plain), 26.2.1911, Millspaugh & Millspaugh 9065 (F); Great Inagua, NW of Morton Bahamas (in open coppice), 23.6.1978, Correll 49939 (US).
Hispaniola: Haiti: Presqu’ile du N.O., Dept. Nord’Ouest, 18 km al este de Mole St Nicolas, en la carretera costera a Jean Rabel (orillas de un río seco en la terraza de terreno donde esta el camino, zona muy arida con Prosopis y Acacia, 19°51'N, 73°17'W, 80 m), 20.1.1984, Zanoni, Pimentel & Garcia 28420 (S); Bombardopolis, south of town in arid mountains (610 m), 25.2.1929, Leonard & Leonard 13549 (US); Tortue Island, vicinity of La Vallee, near harbor, 28.12.1928, Leonard & Leonard 11218 (US); St Louis du Nord, ridge south east of town, 31.3.1929, Leonard & Leonard 14125 (US); Port de Paix, 1 mile NW of city (in dry thicket on hill), 22.12.1928, Leonard & Leonard 11081 (US); Departement du Nord, vicinity of St Michel de l’Atalaye, near the UWJC plantation houses (about 350 m), 17.11.1925, Leonard 7047 (US); Massif du Nord, St Michel, towards La Lome, 500 m, 3.6.1927, Ekman 8340 (US, S); Bayeux, near Port Margot (on sea island, sea level), 9.8.1903, Nash 290 (F); Bayeux, near Port Margot (low land at sea level), 3.8.1903, Nash 30 (F); Borgne (pen hillside, sea level), 14.8.1903, Nash 430 (F); Parish of St Catherine, Mt. Diablo, site of Blue Mountain, 2.5 miles south by road of Hollymount Road (2800-2900 ft), 21.8.1965, Hespenheide & al. 1391 (F). — República Dominicana: Río Orocura 10.1909, Tuerckheim 2573 (BR, F, US, M, S); Prope Fajardo (lit. marino), 12.4.1885, Sintenis 2920 (BR); Province of Santo Domingo, east and north of Howard Allard’s home, 1 mile W of center of city, vicinity of Ciudad Trujillo, waste ground (0.25 m), 5.12.1945, Allard 13896 (US); Province of Santo Domingo, Airport, n.w. of Howard Allard’s home, 1 mi.w. of center of city, vicinity of Ciudad Trujillo, (poor thin sort on coral rock outcrop, waste ground, 0.25 m), 18.11.1945, Allard 13432 (US); Santo Domingo (loam, forests), 7.1921, Faris 348 (US); Santo Domingo (sandy loam, forest tree), 7.1921, Faris 451 (US); Jovero (near sea level), 1.2.1923, Abbott 2559 (US); Jovero (near sea level), 4.12.1923, Abbott 2893 (US); Province of Seibo, La Romana (near the coast), 1.12.1909, Taylor 334 (F); Province of Marcoris, Marcoris River, Consuelo, 22.11.1909, Taylor 261 (F); about 3 miles E of Boca Chica (in thickets), 31.10.1970, Liogier 17657 (F); La Caleta, 20 miles of Santo Domingo City (at sea level), 25.9.1970, Liogier 17501 (F); 1.1871, Wright, Parry & Brummel 304 (F); Provincia Monte Plata, Bayaguana, Carretera a Comatillo, Sabana de los Javalíes (bosque húmedo, alterado, a orilla del camino), 16.6.1997, Jiménez & Veloz 2106A (F); vicinity of Higuey, 3.11.1946, Howard & Howard 9757 (US); Provincia La Romana, Altos de Chavón, ENE of La Romana on upland of W side of Río Chavon (dry arid broadleaf forest and shrubland, open and cut over, 18°25'N, 68°54'W, 75 to 1000 ft), 13.6.1981, Zanoni, Mejía & Ramírez 14861 (US); La Ciénaga (lowland forest), 1100-1200 m, 25.7.1967, Gastony, Jones & Norris 391 (US); Provincia La Romana, La Uvita, S of town of Cumayasa km 10 (on Mar Caribe coast, marine coral limestone, very shallow soil, parts of woody arid forest cut agricultural crops, very dry season now, 18°24'N, 69°03'W, less than 10 m), 24.11.1980, Mejía & Zanoni 9386 (US); Brahona, Santo Domingo, 4.1910, Fuertes 88 (US, BR); Santo Domingo, Ozama, 30.10.1909, Tuerckheim 257b (BR, M, S, US); Santo Domingo, 1.1871, Wright, Perry & Brummel 307 (US), 308 (US); on the top of Diego de Ocampo Peak, 1.1871, Jiménez 4549 (US); La Caleta, 20 miles of Santo Domingo City (at sea level), 25.9.1970, Liogier 17501 (US); 20 km W San Pedro de Macorís (limestone soil), 18.10.1946, Howard & Howard 9469 (FI); Province of Santo Domingo, vicinity of Ciudad Trujillo (0.25 m, waste ground), 5.12.1945, Allard 13896 (S); Province of Santo Domingo, Vicinity of Laguna, Samaná Peninsula, chiefly on the Pilón de Azucarudada, Trujillo (500 m), 18.12.1920, Abbott 301 (US).
Puerto Ric West of Cabo Rojo (200 ft), 20.1.1900, Heller 4421 (US, F); El Verde Ranger Sta. (evergreen seasonal forest, 500-700 ft) 26.6.1950, Little 13110 (F); Camino Cain, near Buena Vista, mountain forest, 1500-2600 ft), 16.7.1950, Little 13411 (F); Camino Hoconuco (dry forest, 1500-2000 ft), 15.7.1950, Little 13389 (F); Rilo 6, Lares Road, near Arecibo, 6.11.1911; Cowles 186 (US); road from Ponce to Adjuntas, 11.7.1901, Underwood & Griggs 770 (US); Isabel Segunda to Campo Rico, 24.1.1914, Shafer 2370 (US); Lares, 400-500 m, 6.4.1913, Britton, Britton & Hess 2749 (US); Martín Peña, 9.8.1914, Stevenson 2185 (US); Puerto Rico, 8.3.1935, Sargent 266 (US); Cayey, south, 15.3.1932, Miller 1612 (US); Pta. Vacía Talega, 2.3 miles NW of Loiza (coastal thicket), 9.8.1989, Lewis & Lewis 14624 (US); Las Croabas, seashore and hill, 2.4.1969, Pfeifer & Class 2920 (US); between Florida and Naguabo, along road, 30.7.1964, Wagner 578 (S); Toa Baja, Sabana Seca, 23.7.1939, Krukoff 681 (F); prope Cayey, 5.10.1885, Sintenis 2321 (US); prope Cabeza de San Juan, 14.3.1885, Sintenis 1915 (S); prope Fajardo (at lit maris versus), 12.4.1885, Sintenis 720 (S); Maunabo, in Punta de la Tuna, 18.9.1886, Sintenis 5088 (S); Cabo Rojo, ad vias ad Joyuda versus, 14.1.1885, Sintenis 720 (S,M); prope Guánica ad salia versus, 2.2.1886, Sintenis 3723 (US); bosque Insular de Guánica (dry forest, 300 ft), 30.6.1950, Little 13176 (F); Hoya Honda Well, Guánica, Mayagüez District (insular forests) 10.10.1940, Gregory 196 (US); east of Ponce, limestone hill, 3.12.1902, Heller (US); Jaburoa (at mare litora), 8.9.1886, Sintenis 4969 (US), Jaburoa (at mare litora), 8.9.1886, Sintenis 4969 (M); prope Fajardo (at maris versus), 12.4.1885, Sintenis 720 (M); N of Ponce, 10.9.1932, Gordon 70 (US); Fajardo and vicitnity, 2.3.1913, Britton & Shafer 1587 (US); Ponce, Guaraguao, 26.11.1899, Goll 827 (US); 3 miles E of Río Piedras, 200 ft, 18.1.1899, Heller 213 (US); Florida (near river), 9.8.1964, Wagner 616 (S); Jabucua (maris litora), 8.9.1886, Sintenis 4969 (F); Fajardo (litora maris versus), 12.4.1885, Sintenis 720b (C); Isla Caja de Muertos, dry forest, alt 50 ft, 21.7.1966, Little 21638 (US); Camino Cain, near Buena Vista (mountain forest), 16.7.1950, Little 13411 (US); Sierra de Luquillo, at kilometer 26, Rt. 191 (on rather open, bushy roadside), 8.9.1967, Hartley 13369 (BR, US); El Verde Ranger Sta., evergreen seasonal forest (500-700 ft), 26.6.1950, Little 13110 (US); St Just, Exp. Forest, 24.8.1950, Little 13741 (F); Laguna Flamenco, 5.6.1968, Woodbury, Martorell & García (F); Palmer, 2 miles N of Mameyes (dry scrubby coastal hills, about 50 m), 6.9.1967, Hartley 1336 (US).
Virgin Islands: St Thomas, 1885, Eggers 90 (M).
Citharexylum spinosum f. smallii (Mold.) I.E. Méndez
Bahamas:Andros, coppice near Deep Creek, Long Bay Cay Sections, 20.-22.1.1910, Small & Carter 8583 (NY [holotype]; Harbor Island, Eleuthera, 18.2.1907, Britton 6402 (F); Mastic Point and vicinity, Andros, 19.3.1907, Brace 7062 (F); Nicholl’s town and vicinity, Andros, 5.3.1907, Brace 6724 (F); Deep Creek, Andros, 18.8.1906, Brace 5140 (F); Mangrove Bay, Andros, 18.9.1906, Brace 4898 (F); Little Creek, S Andros (on copice covered rocky slope), 23.9.1974, Correll 3484 (F); Fresh Creek, Andros, 5.6.1890, Nortrop & Nortrop 608 (F); Eleuthera Point and Vicinity, (rocky scrublands), 24.2.1907, Britton & Millspaugh 5615 (F).
Jamaica: Road to Wareka, 13.1.1904, Harris (US).
Puerto Ric Isla Palominos (dry forest, 50 ft), 2.7.1966, Little 21581 (US).
Citharexylum spinosum f. subserratum (Sw.) I.E. Méndez
Hispaniola: Haiti:Marmelade, 1.8.1905, Nash & Taylor 1328 (US); Ennery (400 m), 27.3.1908, Buch 1089 (S); inter Damasiums & Coteau, 24.6.1917, Ekman H310 (S); vicinity of Bombardopolis, (610 m, dry mountain), 22.2.1929, Leonard & Leonard 13549 (US); inter Constant & Salon, prope Corrail, Dpart. du Sud, 6.9.1917, Ekman H803 (S). — Repùblica Dominicana: Isla Catalina, 1.1950, Vélez 3738 (US).
Citharexylum spinosum f. villosum (Jacq.) I.E. Méndez
Cuba: 1.1941, Howard 4952 (F). — Provincia Habana: Cabana, 24.4.1914, Ekman 297 (S); Cojímar, 12.1906, van Hermann 5069 (F); Cojímar, 7.6.1905, Baber 5137 (F). — Provincia Santa Clara: Pitajones, 28.2.1912, Shafer 12196 (US); Provincia de Camagüey: La Sierrecita, bahía de Nuevas Grandes (bosque semicaducifolio sobre carso), 25.9.1985, Méndez & Trujillo 2066 (HIPC). — Holguín Province: Punta de Piedra, Nipe Bay, 7.3.1912, Britton, Britton & Cowell 12461 (F, US). — Provincia de Santiago de Cuba: El Morro, Santiago de Cuba (in rupibus calcar), 25.9.1916, Ekman 7732 (F, S); Santiago, 21.5.1917, Ekman 8638 (US). — Provincia de Guantánam Imías, 28.5.1982, Bisse & al. 47103 (HAJB, JE). Imías, La Chivera, 20.5.1982, Bisse & al. 48175 (B, HAJB, JE); bosque El Yayal, Limones (bosque siempreverde micrófilo sobre carso), 19.1.1990, Méndez, Caballero & Risco 7629 (HIPC).
Bahamas:Graham’s Harbor, Watling’s Island, 26.11.1907, Wilson 7234 (F); Cockburn Town, Watling’s Island, 25.11.1907, Wilson 7217 (F); Port Hona, Island of Cax, 19.11.1890, Hitchcok (F); New Providence, 11.1890, Hitchcok (F); Cockburn Town and vicinity, Watling’s Island, 12.3.1907, Britton & Millspaugh 6142 (F); Orange Creek and Vicinity, Cat Island, 27.2.1907, Britton & Millspaugh 5744 (F); Jacksonville and vicinity, E Caicos, 26.2.1911, Britton & Millspaugh 9065 (US); San Salvador, N of teachers college and Riding Rock, 21.11.1974, Correll 43855 (F).
Hispaniola: Haiti:Vicinity of Petionville (350 m), 15.1.1920, Leonard 4991 (US); Morne Boutellier, SE of Port au Prince (3000 ft), 11.9.1955, Proctor 10636 (US); Las Cahobas, 28.8.1917, Cook, Scofield & Doyle 89 (US); Petionville (1000 ft), 6.9.1903, Nash 991 (F); Morne Cabrits, Rois d’Ome, Montagnes du Fron d’Eau, 16.7.1924, Ekman H934 (S); Massif de la Selle, Novelle Tronaine, above Chapalle Faure (1400 m), 6.8.1924, Ekman H1372 (S); Morne L’Hôspital, Port au Prince (350 m), 25.8.1942, Holdridge 1428 (S). — Republica Dominicana: Provincia de Monte Cristi, District of La Monción, 30.8.1929, Valeur 112 (US); ibid., 30.5.1931, Valeur 662 (C, F, S, US); Provincia de Monte Cristi, Monción, Cordillera Central (375 m), 28.6.1929, Ekman H13025 (S, US); El Cercado, S of town (750 m, serpentine), 4.9.1968, Liogier 12475 (US); Hondo Valle (pine woodland, 3 500 ft), 1.9.1946, Howard & Howard 8730 (B, S, US).
Puerto Ric Near Aibonito, 28.6.1901, Underwood & Griggs 487 (US); vicinity of Coamo Springs, 2.2.1922, Britton, Britton & Brown 5838 (US); Island of Culebra (rocky hillside), 3.3.1906, Britton & Wheeler 34 (F).
Citharexylum spinosum f. subvillosum (Mold.) I.E. Méndez
Cuba: Wright 3663 (S, US); de la Sagra 207 (B); Romelic Cuba (litoris, 100 m), 4.1889, Eggers 5344 (S, US); El Papayo, Oriente, 22.7.1918, Ekman 9431 (S, US). — Provincia Ciudad de La Habana: Morro de Cojímar 27.4.1914, Ekman 380 (S); Cojímar, 24.8.1910, Britton, Earle & Gager 6301 (US); Playa de Cojímar (scrub or coral limestone near the sea), 16.3.1906, Hitchcock (F); Habana, 6.11.1864, Schott 89 (US); at Río Almendares, 10.11.1921, Ekman 13473 (US); Fortaleza de la Cabaña, 25.9.1904, van Hermann 1958 (F, US); Playa de Marianao, 17.6.1916, León 6255 (US). — Province of Cienfuegos: Cieneguita SW, 14.8.1895, Combs 468 (F); Casa Harward, Soledad, Cienfuegos, González 671 (B). — Provincia Santa Clara: Sierra San Juan, above San Blas (300-600 m), 9.11.1941, Morton 4488 (US); Gavilán, 6.1941, Howard 4952 (US); Provincia de Sancti Spiritus, S de la Sierra del Escambray, Río Cañas (monte seco costero en substrato de piedra caliza, 80°10'W, 21°50'N) 1.7.1993, Acevedo & al. 5542 (US). — Provincia de Camagüey: Cayo Sabinal, costa norte (manigua costera sobre carso), 5.9.1984, Méndez & Trujillo 2677 (HIPC). — Provincia de Las Tunas: Puerdo Padre, Playa Herradura (manigua costera), 6.12.1975, Areces & al. 29028 (B, HAJB). — Provincia Holguín: Preston ad Nipe Bay, viam ferream, 13.11.1914, Ekman 3424 (S, US); Region of Moa, 1.1943, León 21279 (US); El Cayuelo, Guarda la Barca, manigua costera sobre carso 10.10.1985, Méndez & Trujillo 2677 (HIPC). — Provincia Granma: El Guafe, Cabo Cruz, 14.11.1996, Méndez &. Romano 9614 (HIPC). — Provincia Santiago de Cuba: Loma Colorada, 18.4.1914, Ekman 1367 (S); Santiago, austrum versus in pascuis, 21.5.1917, Ekman 8638 (S); Playa de Aguadores, Santiago de Cuba, caminos que bordean la playa, 14.10.1951, Lopez Figueiras 265 (US). — Provincia de Guantánam Guantánamo Bay, 17.-30.3.1909, Britton 2101 (NY [holotype of Citharexylum fruticosum var. subvillosum Mold.]); Baracoa Bay (rup calcareis), 22.11.1914, Ekman 3576 (S); Baracoa, near the lighthouse, 10.1.1915, Ekman 4160 (S); Punta Maisí, 14.12.1910, Shafer 7924 (US).
Jamaica: Long Mountain, (700 ft), 9.11.1904, Harris 8848 (US); Grants Pen, St Thomas, mangrove margin, 16.7.1962, Adams 11364 (M); Galino Point, N of Port Maria (occasional in scrub on pitted limestone flat back of low sea cliff, 2 m), 13.7.1962, Fosberg 42666 (US); St Thomas, Comocan (200 m), 9.1880, Eggers 26 (M).
Bahamas:Orange Creek and vicinity, Cat Island, 27.2.1907, Britton & Millspaugh 5744 (US), 5444 (US); Cat Islands, Hitchcock (F); San Salvador, about 1 mile N of Riding Rock, coppice along roadway through coppice, 13.2.1976, Correll 46717 (F, US); San Salvador, about 1 mile N of Teachers College and Rodong Rock airstrip (coppice), 21.11.1974, Correll 43855 (F); New Providence, Sonetwest Bay (coastal coppice), 3.9.1904, Britton & Brace 499 (F); New Providence, Farrwydon Rod (coppice), 24.8.1904, Britton & Brace 233a (NY [holotype], US [isotype]); Abaco, eight mile Bay, 26.12.1904, Brace 1887 (F); New Providence Island, 2 mil. N of Nassau Airport (secondary forest), 4.11.1982, Little Jr. 37436 (US); 3/4 miles SE of Bottle Creek, N Caicos 11.6.1974, Gillis & Proctor 12237 (B).
Hispaniola: Haiti:Massif de la Selle, nouvelle trousine above Chapelle Faure (1400 m), 6.8.1924, Ekman 1372 (S); Baie des Moustiques, W of Cabaret, Jean Rabal Road (arid thicket) 16.1.1929, Leonard & Leonard 12075 (US). — República Dominicana: Barrabas, 24.4.1906, Raunkiaer 862 (C); Puerto Plata, 27.4.1906, Raunkiaer 1126 (C); Santiago, 3.4.1906, Raunkiaer 878 (C); Provincia de Santiago, Valle del Cibao, Santiago, at El Castillo (hillsides), 4.10.1930, Ekman 16013 (F, S); Province of Monte Cristo, District of Monción (semi-arid pine region), 30.8.1929, Valeur 112 (F, S); Provincia San Cristóbal, Cordillera Central, 2 km al N de Cambita, Garabito, en dieción al Tablazo (finca con Theobroma cacao y arboles frutales, 225 m, 18º27'N, 70º10'W), 6.11.1994, García & al. 5660 (C, F, S); Sto. Domingo, ad López (300 m), 8.6.1887, Eggers 389 (US); El Cercado, Juan Santiago, Hondo Valle (pine woodlands, 3500'), 1.9.1946, Howard & Howard 8730 (FI); Provincia de Barahona, Sal Mine, near La Salina, 16.7.1950, Howard 12068 (B).
Puerto Rico: Island of Culebra (rocky hillside), 3.3.1906, Britton & Wheeler 34 (US); Island of Culebra, Flamingo Bay, 3.3.1906, Britton & Wheeler 215 (F, US); Coamo, roadside, 27.11.1899, Goll, Cook & Collins 695 (US).