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Willdenowia 29 (1/2) - Electronic supplement

Eva Häffner & Frank H. Hellwig

Phylogeny of the tribe Cardueae (Compositae) with emphasis on the subtribe Carduinae:
an analysis based on ITS sequence data

Electronic supplement to:
Häffner, E. & Hellwig, F. H.: Phylogeny of the tribe Cardueae (Compositae) with emphasis
on the subtribe Carduinae: an analysis based on ITS sequence data – Willdenowia 29: 27–39. 1999.

Data matrix

coded gap character                    1
character                             10        20        30        40        50
                              .........|.........|.........|.........|.........|
Ancathia igniaria             TCGAAGCCT--GCACAGCAGAACGACCCGTGAACACGTAATC-ACAACCG
Olgaea pectinata              TCGAAGCCT--GCACAGCAGAACGACCCGTGAACACGTAATC-ACAACCG
Synurus deltoides             TCGAAGCCT--GCATAGCAGAACGACCCGTGAACACGTAATC-ACAACCG
Alfredia cernua               TCGAAGCCT--GCACAGCAGAACGACCCGTGAACACGTAATC-ACAACCG
Onopordum boissieri           TCGAAGCCT--GCACAGCAGAACGACCCGTGAACACGTAATC-ACAACCG
Carlina vulgaris              TCGAAGCCT--GCAAAGCAGAACGACCCGCGAACGTGTAAAACACAACCG
Xeranthemum cylindraceum      TCGAAGCCT--GCACAGCAAAACGACCCGCGAACATGTAACG-ACAATCG
Echinops exaltatus            TCGAAGCCT--GCACAGCAGAACGACCCGTGAACATGTAATG-ACAATCC
Berardia subacaulis           TCGAAGCCT--GCACAGCAGAACGACCCGTGAACACGTAATC-ACAACCG
Stemmacantha rhapontica       TCGAAGCCT--GCACAGCAGAACGACCCGTGAACATGTAATC-ACAATCG
Serratula coronata            TCGAAGCCT--GCACAGCAGAACGACCTGCGAACACGTAATC-ACAACTG
Klasea radiata                TCGAAGCCT--GCACAGCAGAACGACCCGTGAACACGTAATC-ACAACCG
Stephanochilus omphalodes     TCGAAGCCT--GCACAGCAGAACGACCCGTGAACATGTAATC-ATAACCG
Jurinea macrocephala          TCGAAGCCT--GCATAGCAGAACGACCCGTGAACACGTAATC-ACAACAG
Jurinea mollis                TCGAAGCCT--GCATAGCAGAACGACCCGTGAACACGTAATC-ACAACTG
Saussurea riederi             TCGAAGCCT--GCACAGCAGAACGACCCGCGAACACGTAATC-ACAACAG
Cousinia hystrix              TCGAATCCT--GCATAGCAGAACGACCTGTGAACATGTAATC-ACAACCG
Arctium lappa                 TCGAAGCCT--GCACAGCAGAACGACCCGTGAACATGTAATC-ACAACCG
Carduus crispus               TCGAAGCCT--GCACAGCAGAACGACCCGTGGACATGTAATM-ACAGCCG
Carduus nutans                TCGAAGCCT--GCACAGCAGAACGACCCGTGGACATGTAATC-ACAGCCG
Carduus defloratus            TCGAAGCCT--GCACAGCAGAACGACCCGTGGACATGTAATC-ACAGCCG
Cirsium tuberosum             TCGAAGCCT--GCACAGCAGAACGACCCGTGGACACGTAATC-ACAGCCG
Cirsium texanum               TCGAAGCCT--GCACAGCAGAACGACTCGTGGACACGTAATA-ACAGCCG
Tyrimnus leucographus         TCGAAGCCT--GCACAGCAGAACGACCCGTGGACATGTAATC-ACAACAG
Cirsium ciliatum              TCGAAGCCT--GCACAGCAGAACGACCCGCGAACACGTAATC-ACAGCCG
Cirsium eriophorum            TCGAAGCCT--GCACAGCAGAACGACCCGCGAACACGTAATC-ACAGCCG
Notobasis syriaca             TCGAAGCCT--GCACAGCAGAACGACCCGCGAACGCGTAATC-ACA-CCC
Carduus leptacanthus          TTGAAGCCT--GCACAGCAGAACGACCCGTGGACACGTAATC-ACAATCG
Ptilostemon afer              TCGAAGCCTCTGCATAGCAGAACGACCTGTGGACATGTAATC-AATTGTA
Galactites duriaei            TCGAAGCCTCTGCATAGCAGAATGACCTGTGGACATGTAATC-AAATGTA
Cynara humilis                TCGAAGCCT--GCACAGCAGAACGACCCGTGGACAAGTAATC-ACAGCCG
Lamyropsis cynaroides         TCGAAGCCT--GCATCGCAGAACGACCCGTGAACACGTAATC-ACAGCCG
 
coded gap character           2       3              4                  5       
character                             60        70        80        90       100
                              .........|.........|.........|.........|.........|
Ancathia igniaria             RGCGTCGTGGGGATCGAGCGTGAGCTTGAGGCCTGCGATGCTCTGTCGGC
Olgaea pectinata              GGTGTCGTGGGCATCGAGCGTGAGCTCGAGGCCTGCGACGCTCTGTCGGC
Synurus deltoides             GGCGTCGTGGGGATCGAGCGTGAGCTTGAGCCCTGCGATGCTCCGTCGGC
Alfredia cernua               GGCGTTGTGGGCTTCGAGCGTGAGCTCGGGGCCTATGATGCTCCGTCGGC
Onopordum boissieri           GGCGTCGTGGGGATCGGGTGTGAGCTTGAGCCCTGCGACGCTCCGTCGGC
Carlina vulgaris              -GCGGCGTGCGCTTGTGG--------------------------------
Xeranthemum cylindraceum      AGCGACTTGGGGATCGGGCGTTAGCCCGAAACCCTTGAAGCTATGTCGGC
Echinops exaltatus            -GCATCAGGGTGACCGGGTGTGAGCCTGGGAGCTGTGATGCTTTGTTGGT
Berardia subacaulis           GGCGTCGGGGGGATCGGGCGTGA-CGTGAGACC-GCGATGCTCCGCCGGT
Stemmacantha rhapontica       GGTGTCGT-AGGATTGGGTGTGAGCCCTAGCCCTGCGATGCT-CGTCGGC
Serratula coronata            GGTGTCGT-GGGATCGGGTGTGAGCCTGAGCCTTGCGATGCT-CGCCGGC
Klasea radiata                CGTGTCGT-GGGATCGGGTGTGAGCCTGAGCCCTGCGATGCG-CGCCGCC
Stephanochilus omphalodes     GGTGTCGT-GGGATCGGGTGAGAGCCTTAGTCCTACGATGCT-TGCCGAC
Jurinea macrocephala          GGTGTCGCAGGGGTCGGGTGTGAGCCCGGTCCTCGTGATGCTTTGCTGGC
Jurinea mollis                GGTGTTGCGGGGGTCGGGTGTGAGCCTGGTCCTCGTGATGCTTTGCTGGC
Saussurea riederi             GGCGTCTAGGGGATCGGGTGCGAGCCCGAGCCTCGCGATGCTACGCTGGC
Cousinia hystrix              GGCGTCGGGGGGGTCGGGTGCAAGCCTGAGCCTCGTGATGCCCTGTCGGC
Arctium lappa                 GGCATCGGGGGGATTGGGTGCGAGCCTGAGCCTTGCGATGCTTCGTCGGC
Carduus crispus               GGCGTCNAAGGTGTCGGGCGTGAGCCCGGTGCYTGTGATGCCTCGTCGAC
Carduus nutans                GGCGTCAAAGGTGTCGGGCGTGAGCCCGGTGCCTGTGATGCCTTGTCGAC
Carduus defloratus            GGCGTCAAAGGTGTCGGGCGTGAGCCCGGTGCTTGTGATGCCTCGTCGAC
Cirsium tuberosum             GGCGTCGAGGGGGTCGGGCGTGAGCCCGGTGCCYGCGATGYCTCGTCGGC
Cirsium texanum               GGCGTCGAGGGGGTCGGGCGTTA---CG-TGCCCGCGATGCCTTGTCGAC
Tyrimnus leucographus         GGCGTCGAGGGGATCGGGCGTGAGCCTGGTCCTTGTGATGCCCTGCCGAC
Cirsium ciliatum              GGCGTCGGGGGGATCGGGTGTGAGCCCGGTGCCCGCGATGCCTCGTCGAC
Cirsium eriophorum            GGCGTCGGGGGGATCGGGTGTGAGCCCGGTGCCTGCGATGCCTCGTCGAC
Notobasis syriaca             GGCGTTGGGGGGATCAGGCTTGAGCCTGGTGCCCGTGATGCCTCATCGAC
Carduus leptacanthus          GGCGTCGGGGGGATCGGGTGTGAGCCCGGTGTCAGCGATGCCTTGTCGAC
Ptilostemon afer              GGTGTCAGGGGGATTGGGTGTGAGCCCGATGCCTGTGATGCCTTGCTGGT
Galactites duriaei            RGTGTAATGGGGATCAGGTGTGAACCTGATTCCTGTGATGCCTTGCTGGT
Cynara humilis                GGCGTCGAGGGGGTCGGGTGTGAGCCCGATGCTCACGATGCCCCGTCGAC
Lamyropsis cynaroides         GGCTTCGGAGGGATTGGGCGTGAGCCTGTTTCCTGCAAAGCCTCGTCGGC
 
coded gap character                             6          7          89 10     
character                            110       120       130       140       150
                              .........|.........|.........|.........|.........|
Ancathia igniaria             GTGCGTTTAGGGTGCCCC-TTTCG-AGGCATCATGGGCGTCAT-GTCGTC
Olgaea pectinata              GAGCGTTCAGGGTGCCTT-TTTTG-AGGCATCGTGGGCGTC-TTGTCGTC
Synurus deltoides             GTGCGTTCAGGGTGCCCC-TTTCG-AGGCATCATGGGCGTCGTCGTCGTC
Alfredia cernua               GTACGTTCAGG-TGCCCT-TATTG-AGGCATCATGGGCGTC-TCGTCGTC
Onopordum boissieri           ATGCGTTCAAGATGCCCC-TTCCGGAGGCATCATGGGCGT-ATCGTCGTC
Carlina vulgaris              --------------CCTC-CCTTT-GGGTGCCTCGGGTGTT-TCGTCGCC
Xeranthemum cylindraceum      GTACGTGTGCTGTTCCAT-CTTTG-GAAAGTAGCATACGT-GC-GTTGGC
Echinops exaltatus            GTGCGCACACCGGGTCAC-TTTGT-GGCC-TTGTGGATGTTAT-GCCGAC
Berardia subacaulis           GTGCGTGCACTGTGCCTC-TTTTG-AGCGATCGTGGACGTTGC-ATCGGC
Stemmacantha rhapontica       ATGCGTCCAAGGTGCCTA-TCTTT-AGGCATCGTGGATGTT-TTGTCGGC
Serratula coronata            ATGCGTGCAAGGTGCCTA-TCTTT-TGGCATCGTGGATGTTGT-GTCGGC
Klasea radiata                ATGCGTGCAAGGTGCCTA-TCTTT-AGGCATCGAGGACGTTGT-GTCGGC
Stephanochilus omphalodes     ATGCGTGCAAGGTGCCTA-TATGT-AGGTGCCGTGGACGTTGT-GTCGGC
Jurinea macrocephala          ATGCATGCATGGTGTTTC-TTGCG-AGGCACTGTGTGTGTTGT-GTTGGC
Jurinea mollis                ATGCATGCATGGTGTTTC-TTGCG-AGGCACCGTGGGTGTTGT-GTTGGC
Saussurea riederi             ATGCGTGCATGGTGTCCC-TTTTG-AGGCATCATGGACGTCGT-GTCGGC
Cousinia hystrix              GTACGTGCATGGTTCCCC-TTATA-AGGCATCGTGGACGTTGC-GTTGGC
Arctium lappa                 ATACGTGCATGGTGCCCC-TTACG-AGGCATTGTGGATGTTGT-GTCGGC
Carduus crispus               GTGTCTCTGCGTTGCCCCGTTTTG-GGGCGTCGTGGATGTYGC-GTYGGV
Carduus nutans                GTGTCTCTGCGTTGCCCCGTTTTG-GGGCGTCGTGGATGTTGC-GTCGGC
Carduus defloratus            GTGTCTCTGCGTTGCCCCGTTTTG-GGGCGTCGTGGATGTCGC-GTCGGC
Cirsium tuberosum             GTGCGTCTGCGATGCCCCGTTTCG-AGGCGTCGTGGATGTTGC-GTCGGC
Cirsium texanum               GTGCGTCCGCGATGCCCCGTTTCG-CGGT-CTGTGGATGTTGC-GTCGGC
Tyrimnus leucographus         GTGGGTCAGCAATGCCTTGTTTCA-AGGCGTCGTGGATTTTGC-GTCGGC
Cirsium ciliatum              GTGCGTCCGCAATGCCCCGTTTCG-AGGCATCGTGGATGTTGT-GTCGGC
Cirsium eriophorum            GTGCGTCCGCAATGCCCCGTTTCG-AGGCATCGTGGATGTTGT-GTCGGC
Notobasis syriaca             GTGCGTCTGCGATGCCTCGTTTCG-AGGAATCGTGGATGTTGT-GTCGGC
Carduus leptacanthus          GTACGTTCGTGATGCCCCGATTCG-AGGCGTCGTGGATGTTGC-GTTGGC
Ptilostemon afer              ATGCGTGCATGGTGCCCCGATTCG-AGGCATTGTGGATGTCAT-GTCGGC
Galactites duriaei            ATGCGTGCATGGTGCCTCGATTTG-AGGCATTGTGGATGTCAT-GTCGGC
Cynara humilis                GTGCGTGCATGGTGCCCCGTTTTG-AGGCATCGTGGACGTCGC-GTTGGC
Lamyropsis cynaroides         GTGCGTTCATGTTGC-TCGTAACG-AGGCATCATGGATGTTGC-GTCGGC
 
coded gap character                      11         12      13                  
character                            160       170       180       190       200
                              .........|.........|.........|.........|.........|
Ancathia igniaria             ACCGAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACCAAATTTAAG
Olgaea pectinata              ACCGAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACCAAATTTAAG
Synurus deltoides             ACCGAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACCAAACTTAAG
Alfredia cernua               ACCGAAACAAAACCCCGGCACGGCATGTGCCAAGGAAAACAAAACTTAAG
Onopordum boissieri           ACCGAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACTTAAG
Carlina vulgaris              ACGTAAACAAA-CCCCGGCACAACACGTGCCAAGGAAAACAAAACTTAAG
Xeranthemum cylindraceum      CCAAAAACCAA-CCCCGGCACGGCATGTGCCAAGGAAATAAAAACTTAAG
Echinops exaltatus            ACAAAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACATAAG
Berardia subacaulis           ACGAAAACAAA-CCCCGGCACG-CATGTGCCAAGGAAAACCAAACTTAAG
Stemmacantha rhapontica       ACAAAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACTTAAG
Serratula coronata            ACAAAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACTTAAG
Klasea radiata                ACAAAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACTTAAG
Stephanochilus omphalodes     GCATAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAACCTTAAG
Jurinea macrocephala          ACGAAAACAAACCCCCGGCACGGCATGTGCCAAGGAAAACAAAACATAAG
Jurinea mollis                ATGATAACAAACCCCCGGCACGGCATGTGCCAAGGAAAACAAAACATAAG
Saussurea riederi             ATGAAAACAAACCCCCGGCACGGCATGTGCCAAGGAAAACCAAACTTAAG
Cousinia hystrix              ACAAAAACAAAACCCCGGCACGGTATGTGCCAAGGAAAACAAAACTCAAG
Arctium lappa                 ACGAAAACAAA-CCCCGGCACG-CATGTGC-AAGGAAAACCAAACTTAAG
Carduus crispus               ACTTWAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACATAGG
Carduus nutans                ACTTTAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACATAGG
Carduus defloratus            ACCTAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACATAGG
Cirsium tuberosum             ACCTAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACCTAGG
Cirsium texanum               ATCTAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACATAGG
Tyrimnus leucographus         ACCTAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACATAAG
Cirsium ciliatum              ACCTAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACATAAG
Cirsium eriophorum            ACCTAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACATAAG
Notobasis syriaca             ACCTAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACATAAG
Carduus leptacanthus          ATCTAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACAAAACATAAG
Ptilostemon afer              ACTTAAACAAA-CCCCGGCACGGCAAGTGCCAAGGAAAATCAAACAAAAG
Galactites duriaei            ACCTAAACAAA-CCCCGGCACAGCAAGTGC-AAGGAAAATCAAACAAAAG
Cynara humilis                ACGTAAACAAA-CCCCGGCACG-CATGTGC-AAGGAAAACAAAACATTAG
Lamyropsis cynaroides         ACGTAAACAAA-CCCCGGCACGGCATGTGCCAAGGAAAACGAAATATAAG
 
coded gap character                                                        14   
character                            210       220       230       240       250
                              .........|.........|.........|.........|.........|
Ancathia igniaria             AAGGGTRCGT-CTCGTGTAGCCCC-GTTCGCGGTGTGCGCACGTGTCGTG
Olgaea pectinata              AAGGGCACGT-CTCGTGTTGCCCC-GTTCGCGGTGTGCGCATGTGTCGTG
Synurus deltoides             AAGGGCGCGT-CTCGTGTTGTCCC-GTTCGCGGTGTGCGCACGTGTCGTG
Alfredia cernua               AAGGGCGCGT-CTCGTGTTGCCCC-GTTCGCGGTGTGCGCGCGTGTCGTG
Onopordum boissieri           AAGGGCGCGT-CTCGTGTTGCCCC-GTCCGCGGTGTGCGCACGGGCCGTG
Carlina vulgaris              AAGGGCGCGT-CCCGTGTTGCCCC-GTTCGCGG-GAGCGCACGGG-CGTG
Xeranthemum cylindraceum      ATGGGCGCGT-CCGGTGTCGCCCC-GTTCTCGGTGTGCGCACAGG-CGTG
Echinops exaltatus            AAGGGTGCA--CCCGTGTTGCTCC-GTTCGCGGTATGCGCATGGGTCGTG
Berardia subacaulis           AAGGGCGCGT-CCCGTGTTGCCCC-GTTTGCGGTATGCGCATGGGCCGTG
Stemmacantha rhapontica       AAGGGTGCGT-CTCGTGTTGTCCC-GTTTTCGGTGCGCACACAGGTCGTG
Serratula coronata            AAGGGTGCGT-CTTGTGTTGCCCT-GTTTTCGGTGTGCACACGGGTCGTG
Klasea radiata                AAGGGTGCGT-CTCGTGTTGCCCC-GTTTTCGGTGTGCACATGGGTCGTG
Stephanochilus omphalodes     AAGGGTGCGT-CACGTGTTGTCCC-GTTTTCGGTGTGCACACGGGTTGTG
Jurinea macrocephala          AAGGGTGCGT-CCCGTGTTGCCCC-GTTTGCGGTGTGCGCATGGGTCGTG
Jurinea mollis                AATGGCGCGT-CCCGTGTTGCCCC-GTTCGCGGTGTGCGCGTGGGTCGTG
Saussurea riederi             AAGGGCGCGG-CTCGTGTTGCCCC-GTTTGCGGTGTGCACATGGGTCGTG
Cousinia hystrix              AAGGGCGCGT-CCTGTGTCGCCTCCGTTTGCGGTGTGCGCATGGGTCGTG
Arctium lappa                 AAGGGCGCGT-CCTGTGTTGCCCC-GTTTGCGGTGTGCGCACGGGTCGTG
Carduus crispus               AAGGACGCGT-CTCGTGMTGCCTC-GTTCGCGGGGTGKGCAYGGGTCGTG
Carduus nutans                AAGGACGCGT-CTCGTGCTGCCTC-GTTCGCGGGGTGGGCACGGGTCGTG
Carduus defloratus            AAGGTCGCGT-CTCGTGATGCCTC-GTTCGCGGGGTGTGCATGGGTCGTG
Cirsium tuberosum             AAGGGCGCGT-CCCGTGTTGCCCC-GTTCGCGGTGTGYGCACGGGTCGTG
Cirsium texanum               AAGGGCGCGT-CCCGTGTTGCCCC-GTTCGCGGTGTGCGCATGGGCCGTG
Tyrimnus leucographus         AAGGGCTTGT-CTCGTGTTGCCCC-GTTCGCGGTGTGCGCATGGGTCGTG
Cirsium ciliatum              TAGGACGCGT-CCCGTGTCGCCCC-GTTCGCGGTGTGCGCACGGGCCGTG
Cirsium eriophorum            TAGGACGCGT-CCCGTGTCGCCCC-GTTCGCGGTGTGCGCACGGGCCGTG
Notobasis syriaca             AAGGGCGCGT-CTCGTGTTGCCCC-GTTTGCGGGGTGTGCACGGGCGATG
Carduus leptacanthus          AAGGGCGCGT-CTCGTGTAGCCTC-GTTCGCGGTGTGCACACGGGTCGTG
Ptilostemon afer              AAGGTCGTGT-CCCGTGTTGCCCC-GTTCGCGGTGTTTGCATGGGCTATG
Galactites duriaei            AAGGTCGTGT-CCCGTGTTGCCCC-GTTCGTGGTGTTTGCATGGGCTGCG
Cynara humilis                AAGGGCGCGTGCTCGTGTCAC-TC-GTTCGCGGAGTGCAC-CGGCCCGTG
Lamyropsis cynaroides         AAGGGTGCGT-CCTTTGTTGCCCC-GTTTGCGGTGTGCGCATGGGTCGTG
 
coded gap character                                                 18          
coded gap character             15        16                       1719   2021  
character                            260       270       280       290       300
                              .........|...ITS1ITS2.. .....|.........|.........|
Ancathia igniaria             GCCTCTCATTAA--CCA?????????????????????????????????
Olgaea pectinata              GCCTCTCATTAAAATAAATCGCGTC-GCCTCCGACCTCGCCTCTCCCATG
Synurus deltoides             GCCTCTCATTAA--CCAATCGCGTC-GCCTCCGACCACGCCTCTCCCATA
Alfredia cernua               GCCTCTCATTAA--CCA?????????????????????????????????
Onopordum boissieri           GCCTCTCATTAA--CCAATCGCGTC-GCCTCCGACCACGCCTCTCCCATG
Carlina vulgaris              GC-TCTCTAGAA--CCAATCGCGTC-GCTCCCGACCACGCCTCCCCCACG
Xeranthemum cylindraceum      GCCTCTTTTTAA--CCAATCGCGTC-GCTCCCGACCGC-TCTCC-CTTCG
Echinops exaltatus            GCCTCTTTGAAA--CCAATCGCGTC-GCCCCTGACCAT-CATCC--TATG
Berardia subacaulis           GCTTCTCAATAA--CCAATCGCGTC-GCCTCCGACCACGCCTCCCCATTG
Stemmacantha rhapontica       GCCTCTCATTAA--CCAATCGCGTC-GCCCCAGACCACG-CTCCCC-ATG
Serratula coronata            GCCACTCAATAA--CCAATCGCGTC-GCCCTAGACCAC-CCTCCCTCATG
Klasea radiata                GCCTCTCAATAA--CCAATCGCGTC-GCCCCAGACCATG-CTCCCCTAAG
Stephanochilus omphalodes     GCCTCTCATTAA--CCAATCGCGTC-GCCCCAGACCAT-TCTCCCCCATG
Jurinea macrocephala          TCTTCTCATTAA--CCAATCGCGTC-GCGCCCCACCACGCCTCCCTTATG
Jurinea mollis                TCTTCTCATAAAA-CCAATCGCGTC-GCCCCCGACCACGCCTCCCTCATG
Saussurea riederi             GTCTCTCATTAA--CCAATCGCGTC-GCCCCCAACCACGCCTCCCTCATG
Cousinia hystrix              GCCACTCACCAA--CCAATCGCGTC-GCCCCCGACCA-GCCTCCCCTGTG
Arctium lappa                 GCCTCTCATCAAA-CCAATCGCGTC-GCCCCCGACCA--CCTCCCA-GTG
Carduus crispus               GCCTCTCAATAA--CCAATCGCGTG-GCCCCAGACCACGCCTCC-ATATG
Carduus nutans                GCCTCTYAATAA--CCAATCGCGTC-GCCCCAGACCACGCCTCC-ATATG
Carduus defloratus            GCCTCTCAATAA--CCAATCGCGTC-GCCCCAGACCACGCCTCC-ATACG
Cirsium tuberosum             GCCTCTCAATAA--CCAATCGCGTC-GCCCCAGACCACGCCTCCCCAACG
Cirsium texanum               GC-TCTCGATAA--CCAATCGCGTC-GCTCCCGACCACGCCTGCCC-ATG
Tyrimnus leucographus         GCCTCTCAATAA--CCAATCGCGTC-GCCCCAGACCTCGCCTCCCCAACG
Cirsium ciliatum              GCCTCTCAATAA--CCAATCGCGTCAGCCCCAGACCACGCCTCCCCAACG
Cirsium eriophorum            GCCTCTCAATAA--CCAATCGCGTC-GCCCCAGACCATGCCTCCCCAACG
Notobasis syriaca             GTCCCTCAATAA--CCAATCGCGTC-GCCCTGGACCATGCCTCCCTAAAG
Carduus leptacanthus          GCCTCTCAATAA--CCAATCGCGTC-RCCTCAGACAACGTCTTCCCAATG
Ptilostemon afer              GCCTCTCATTAA--CCAATCGCGTC-GCCCCAGACTATGTCTCCTCTATA
Galactites duriaei            GCCTCTCATTAAA-CATATCGCATC-GCCCCAGAT--TGTCTCCTCTATA
Cynara humilis                GC-TCTCAAAAA--CCAATCGCGTC-GCCCCAGACCACGACTCCTCGATG
Lamyropsis cynaroides         ACCTCTCAATAA--CCAATCGCGTC-GCCCCAGATCAWGCACCCCCGATG
 
coded gap character             23        28  30                                
coded gap character            22 2425 2627  293132   33  34                    
character                            310       320       330       340       350
                              .........|.........|.........|.........|.........|
Ancathia igniaria             ?????T-GCGTT-TG-GTC-GGGGG-CGGAGATTGGTCTCCCGTGTCCAT
Olgaea pectinata              GGG-AT-GCGTT-TG-G-CCGGGGG-CGGAGATTGGTCTCCTGTGTCCAT
Synurus deltoides             GGG-AT-GCGTT-TG-GTCCGGGG--CGGAGATTGGTCTCCTGTGTCCAT
Alfredia cernua               ??????????????????????????????????????????????????
Onopordum boissieri           GGG-AC-GCGTT-TG-GTCCGGGGG-CTGAGACTGGTCTCCCGTGCCCAT
Carlina vulgaris              GGG--TGGAGTGCCG--TC-GGGGG-CGGAGACTGGCCTCCCGTGCCCAT
Xeranthemum cylindraceum      G---AC--TGTTTTG--TC-GTGTG-CGGATATTGGTCTCCCGTTCCGAT
Echinops exaltatus            G---AC-AAGTG--GTGTC-AGGGAGCGGATATTGGTCTCCCGTGCTCAT
Berardia subacaulis           GG--AT-GTGTTTTG--TC-GGGGA-CGGAGACTGGTCTCCCGTGCCCAT
Stemmacantha rhapontica       GG--AT-GTGTTTTA--TCTGGGA--CGGAGACTGGTCTCCCGTGCCGAC
Serratula coronata            GGG-AT-GTGTTTCG--TCTAGGA--CGGAGACTGGTCTCCCGTGCTGAA
Klasea radiata                GG--AT-GTGTTTTG--TCTGGGA--CGGAGACTGGTCTCCCGTGCCGAT
Stephanochilus omphalodes     G---AC-GTGTT-TG-GTTTGGGA--CGGAGACTGGTCTCCCGTTCCGAT
Jurinea macrocephala          GGG-AT-GGGT--TGTGTC-GGGGG-CGGAGACTGGTCTCCCGTGCCCAT
Jurinea mollis                GGA-AT-GGCT--TGTGTC-GGGA--CGGAGACTGGCCTCCCATGCCCAT
Saussurea riederi             GGG-AC-GTGTTTTG--TTTGGGG--CGGAGAATGGTCTCCCGTGCTCAT
Cousinia hystrix              GGG-AT-GCGTCTCG--TCCGGGG--CGGAGACTGGCCTCCCGTGCCCAT
Arctium lappa                 GGG-ATTGCGTGTCG--TTTGGGG--CG-AGACTGGTCTCCCGTGCCCAT
Carduus crispus               GGGGAT-GTGTT-TG--TCTGGGG--CGGAGAATGGTCTCCCGTGCCGTC
Carduus nutans                GGGGAT-GTGTT-TG---CTGGGG--CGGAGAATGGTCTCCCGTGCCGTT
Carduus defloratus            GGGGAT-GTGTA-TG--TC-GGGG--CGGAGAATGGTCTCCCGTGCCGTC
Cirsium tuberosum             GG--AT-GCGTT-CG--TCYGGGG--CGGAGAATGGTCTCCCGTGC-GTC
Cirsium texanum               GGG-AT-GCGTT-CG--TCTGGGG--CGGAGAATGGTCTCCCGTGCCGTC
Tyrimnus leucographus         GG--AT-GC-TTTCG--TCTGGGG--CGGAGAATGGTCTCCCGTGCCGTC
Cirsium ciliatum              GG--AT-GG-TT-CG--TT-GGAG--CGGAGAATGG-CTCCCGTGCCGTT
Cirsium eriophorum            GG--AT-GCG-CTCG--T--GGAG--CGGAGAATGGTCTCCCGTGCCGTT
Notobasis syriaca             TGG-AT-GCGTT-CG--TCTGGGG--CGGAGAATGGTCTCCCGTGCCGGT
Carduus leptacanthus          GGG-AT-CCGTT-CA--TCTGGGG--CGGAGAATGGTCTCCCGTGCCGTT
Ptilostemon afer              GGG-AC-GTGTTTTG--TCTGGGG--CGGATACTGGTCTCCCGTGCTTCT
Galactites duriaei            GGG-AC-GTGTTTTG--TCTGGGG--CG-AAAATGGTCTCCCGTGCTTTT
Cynara humilis                GGG-ATTCGGTTTTG--TCTGGGG--CG-AGACTGGTCTCCCGTGCCTTT
Lamyropsis cynaroides         GG--AT-GCGTT-AG-GTCTGGGT--CGG-GATTGGTCTCCCGTGTCGTT
 
coded gap character              35 36                   37                     
character                            360       370       380       390       400
                              .........|.........|.........|.........|.........|
Ancathia igniaria             GGCGCGGTTGGCCTAAAACGGAGTCCCC-TCCGATGGACGCA-CGGCTAG
Olgaea pectinata              GGCGCGGTTGGCCTAAAACAGAGTCCCC-TCCGATGGACGCA-TGGCTAG
Synurus deltoides             GGCGCGGTTGGCCTAAAACGGAGTCCCC-TCCGATGGACGCA-TGGCTAG
Alfredia cernua               ??????????????????????????????????????????????????
Onopordum boissieri           GGC-CGGTTGGCCTAAAAAGGAGTCCCC-TCCGACGGACGCA-CGGCTAG
Carlina vulgaris              GGCGCGGCTGGCCCAAACAGGAGTCCCC-TTCGACGGACGCA-CGACTAG
Xeranthemum cylindraceum      GGAGCGGTTGGCCTAAAAAAGAGTTCCC-TCCGGCAAACGCA-TGGCTAG
Echinops exaltatus            GGTGTGGTTGATCTAAATAGGAGTCCTC-TTCGGTGGATGCA-CGACTAG
Berardia subacaulis           GGCGCGGTTGGCCTAAAAAGGAGTCCCCCTTCGACGGACGCA-CGGCTAG
Stemmacantha rhapontica       GGCGCGGTTGGCCTAAAAAGGAGTCCCC-TTTGGCGGGCGCA-CGGCTAG
Serratula coronata            GGCGCGGTTGGCCTAAAAAGGAGTCCCC-TTTGGCGGACGCA-CGGCTAG
Klasea radiata                GGTGCGGTTGGCCTAAAAAGGAGTCCCC-TTTGGCGGACGCA-CGGCTAG
Stephanochilus omphalodes     GGTGCGGTTGGCCTAAAAAGGAGTCCCC-TTTGGCGGACGCA-CGGCTAG
Jurinea macrocephala          GGTGTGGTTGGCCTAAAAAGGAGTCCCC-TTTGACGGACGCA-CGACTAG
Jurinea mollis                GGTGTGGTTGGCCTAAAAAGGAGTCCCC-TTTGTCGGACGCA-CGGCTAG
Saussurea riederi             GGTGCGGTTGGCCTAAAAAGGAGTCCCC-TTCGACGGACGCA-CGGCTAG
Cousinia hystrix              GGTGCGGTTGGCCTAAAAAGGAGTCCCC-TTTGACGGACGCA-CGGCTAG
Arctium lappa                 GGTGCGGTTGGCCTAAAAAGGAGTCCCC-TTTGACGGACGCA-CGGCTAG
Carduus crispus               GGCGCGGTTGGCCTAAAAAGGAGTCCCC-TTCGACGGACGCA-CGGCTAG
Carduus nutans                GGCGCGGTTGGCCTAAAAAGGAGTCCCC-TTCGACGGAGGCA-CGGCTAG
Carduus defloratus            GGCGCG-TTGGCCTAAAAAGGAGTCCCC-TTCGACGGATGCA-CGGCTAG
Cirsium tuberosum             GGCGCGGTTGGCCTAAAAAGGAATCCCC-TTCGACGGACGCA-CGGCTAG
Cirsium texanum               GGCGCGGTTGGCCTAAAAAGGAGTCCCC-TTCGACGGACGCA-CGGCTAG
Tyrimnus leucographus         GGCGCGGTTGGCCTAAAAAAGGAGTCCC-GGCGACGGACGCA-CGGCTAG
Cirsium ciliatum              GGCGTGGTTGGCCTAAAAAGGAGTCCCC-TTCGA-GGACGCA-CGGCTAG
Cirsium eriophorum            GGCGTGGTTGGCCTAAAAAGGAGTCCCC-TTCGACGGACGCA-CGGCTAG
Notobasis syriaca             GGC----TTGGCCTAAAAAGGAGTCCCC-TTTGACGGGCGCA-CGGCTAG
Carduus leptacanthus          GGTGTGGTTGGCCTAAAAAGGAGTCCCC-TTCGACGGACGCA-CGGCAAG
Ptilostemon afer              GGTGCGGTTGGCCTAAAAAGGAGTCCCT-TTTGACGGACGCA-CGGTTAG
Galactites duriaei            GGTGCGGTTGGCCTAAAAAGCAGTCCCT-TTTGTCGGATGCA-CGGCTAG
Cynara humilis                GRTGCGGTTGGCCTAAAAAGGAGTCCC--TTCGGCGGATGCA-CGGCTAG
Lamyropsis cynaroides         GGCGTGGTTGGCCTAAAAAGGAGTACC--TTCGACGGACGCAACGACTAG
 
coded gap character                           38                39  40 41    42 
character                            410       420       430       440       450
                              .........|.........|.........|.........|.........|
Ancathia igniaria             TGGTGGTTGTCAAGGC-TTCGTATCGAGCCGTGCGTCGTGAGTCGTG-AG
Olgaea pectinata              TGGTGGTTGTCAAGGC-TTCGTATCGAGCCGTGCGTCGT-AGTCGCG-AG
Synurus deltoides             TGGTGGTTGTCAAGACCTTCGTATCGAGCCGTGCGTTGTGAGTCGTG-AG
Alfredia cernua               ??????????????????????????????????????????????????
Onopordum boissieri           TGGTGGTTGTCAAGGCCTTCGTATCGAGCCGTGCGTCGTGAGCTGCG-AG
Carlina vulgaris              TGGTGGTTGTAACGGCCCTCGTATCGAGCCGTGCGTCTTGAGCCGCA-AG
Xeranthemum cylindraceum      TGGTGGTTGTAATGGCCTTCGTATCGAGTCGTGTGTTGTGAGCCGCA-AG
Echinops exaltatus            TGGTGGTTGTATAATC--TCGTATCGAGCCGTGTGTTGTGAGCCGCA-AG
Berardia subacaulis           TGGTGGTTGTCAAGGCCTTCGTATCGAGCCGTGTGTCGTGAGCCGCA-AG
Stemmacantha rhapontica       TGGTGGTTGTCAAGGCCTTCGTATCGAGCCGTGT----TGA--TGCA-AG
Serratula coronata            TGGTGGTTGTCAAGGCCTTCGTATCGAGCCGTGC----TGA--AGCA-AG
Klasea radiata                TGGTGGTTGTCAAGGCCTTCGTATCGAGCCGTGT----TGA--TGCA-AG
Stephanochilus omphalodes     TGGTGGTTGTCAAAGCCTTCGTATCGAGTCGTGC----TGA--TGCT-AG
Jurinea macrocephala          TGGTGGTTGTCGAGACCTTTGTAACGAGCCGTGC----GGA--TGCAAAG
Jurinea mollis                TGGTGGTTGTCAAGGCCTTCGTAACGAGCCATGT-----GA--TGCAAAG
Saussurea riederi             TGGTGGTTTTCAAGGCCTTCGTATCGAGCTGTGC----ATA--CGCG-AG
Cousinia hystrix              TGGTGGTTGTCAAGGCCTTCGTATCGAGCCGTGC----GGA--CGCA-AG
Arctium lappa                 TGGTGGTTGTCAAGGCCTTCGTATCGAGCCGTGC-----GA--CGCA-AG
Carduus crispus               TGGTGGTTGAAAA-GCCTTCGTATCGAGCCGTGTGTCGTTAGCCGCATAG
Carduus nutans                TGGTGGTTGAAAAAGCCTTCGTATCGAGCCGTGTGTCGTTAGCCGCA-AG
Carduus defloratus            TGGTGGTTGATAAGGCCTTCGTATCGAGCCGTGTGTCGTTAGCCGCA-AG
Cirsium tuberosum             TGGTGGTTGTTAAGGCCTTCGTATCGAGCCGTGTGTCGTTAGCCGCA-AG
Cirsium texanum               TGGTGGTTGTTAAGGCCTTCGTATCGAGCCGTGCGTCGTTAGCCGCA-AG
Tyrimnus leucographus         TGGTGGTTGATAAGTCCTTCGTCTCGAGCCATGTGTCGTTAGCCGCA-AG
Cirsium ciliatum              TGGTGGTTGTTAAGGCCTTCGTATCGAGTCGTGCGTCGTTACGCGCA-AG
Cirsium eriophorum            TGGTGGTTGTTAAGGCCDG-GTATCGAGTCGTGCGTCGTTAGCCGCA-AG
Notobasis syriaca             TGGTGGTTGTTAAGGCCTTCGTATCGAGCCGTGGGTTGTTAGTCGCA-AG
Carduus leptacanthus          TGG---TTGTTAAGGCCTTCGTATTGAGCTGTGCGTCGTGAGCCGCA-AG
Ptilostemon afer              TGGTGGTTGTCAATGCCTTCGTATTGAGCCGTGTGTCTTGATCCATA-AG
Galactites duriaei            TGGTGGTTGTCAAGGCCTTCGTATTGAGCCGTGTGTCGTGATCTATA-AG
Cynara humilis                TGGTGGTTGTCAAGGCCTTCGTATCGAGCCGTG-GTCGGGTGCCGCG-AG
Lamyropsis cynaroides         TGGAGGTTTTCAAGGACTTCGTATCGAGTCGTACGCCGTGAGTCGTA-AG
 
coded gap character                                         44                  
coded gap character                                        4345            46   
character                            460       470       480       490       500
                              .........|.........|.........|.........|.........|
Ancathia igniaria             GTAAATGCTCTCCAAAGACCCCATTG-TGTC--CTTGCGACGATGTTTCG
Olgaea pectinata              GTAAACGCTCTCCAAAGACCCCAATG-TGTCGTCTTGTGACGATGCTTCG
Synurus deltoides             GTAAACGCTCTCCAAAGACCCCAATG-TGTCGTCTTGTGACGATGCTTCG
Alfredia cernua               ??????????????????????????????????????????????????
Onopordum boissieri           GGAAGCGCTCTCCAAAGACCCCAACG-CGTCGTCTTGCGACGATGCTTCG
Carlina vulgaris              GGAAGCGCTCGACAAAGACCCCAATG-CG-CGCCTCGCGACGACGCTTCG
Xeranthemum cylindraceum      GGAATTGCTCTTTAAAGACCTTTATG-CGTCGTCTGTTGACGACGTTTCG
Echinops exaltatus            GGAGTTTCTCTTCAAAGACCCCTTAG-TGTCGTCTTGTGACGATGCTTCG
Berardia subacaulis           GGAAGCGCTCTCCAAAGACCCCAACA-CGT-GTCTTGCGACGATGCTTCG
Stemmacantha rhapontica       GGATTTGCTCTCTAAAGACCCTAACG-TGTCGTCTTACGACGATGCTTCG
Serratula coronata            GGAATCGCTCTTTAAAGACCCTAACG-TGTTGTCTTACGACGATGCTTCG
Klasea radiata                GGAATCGCTCTCTAAAGACCCCAACG-TGT-GTCTTGCGACGATGCTTCG
Stephanochilus omphalodes     GGAATCGCTCTCTAAAGACCCTAATG-TGTCGTTTTACGACGATGCTTCG
Jurinea macrocephala          GGATACGCTCTATAAAGATCCCAACG-TGTCGTCTTGTGACAATGCTTCG
Jurinea mollis                GGAT-CGCTCTAAAAAGACCCCAACG-TGCTGTCTTATGACGACACTTCG
Saussurea riederi             GGATTCGCTCTCCAAAGACCCCAACG-CGTCGTCTTGCGACGACGCTTCG
Cousinia hystrix              G-AAGCGCTCTCCAAAGACCCCAA----GCCRTCTCGTGACGAC??????
Arctium lappa                 GGAAGCGCTCTCCAATGACCCCAACG-TGTCGTCTTGCAACGACGCTTCG
Carduus crispus               GGAAGCGCTCTCCATAGACCCTAACG-TGTC--CTCGCGACGA-GCAACG
Carduus nutans                GGAAGCGCTCTACATAGACCCTAACG-TG-C--CTCGCGACGATGCAACG
Carduus defloratus            GGAAGCANTCTCCGTAGACCCTAATG-TGTCGTCTCGTGACGATGCTTCG
Cirsium tuberosum             GGAAGCGCTCTCTAAAGACCCCAACG-C-TCGTCTYGCGACGAYG-TTCG
Cirsium texanum               GGAAGCGCTCTCTAAAGACCCCAACG-TGTCGTCTCGCGACGACGCTTCG
Tyrimnus leucographus         GGATGCGCTCTCCGAAGACCCCAACGGTGTTTTCTTGCGACGACTCTTCG
Cirsium ciliatum              GGAAGCGCTCTCCAAAGACCCTAACG-TGTGTTCTAGCGACGATGCTTCG
Cirsium eriophorum            GGAAGCGCTCTCCAAAGACCCTAACG-TG---TCTAGCGACGATG-TTCG
Notobasis syriaca             GGAAGTGCTCCCCAAAGATCCCAACG-TGTCGAATAGTGACGATGCTTCG
Carduus leptacanthus          GGAAGCG-TCTCCAAAGACCCCAACG-TGTCGTCTTGTGACGATGCTTCG
Ptilostemon afer              GGAAGCGCACTCCAAATACCTCAATG-TGTCGTCTCTTGACGACACTTCG
Galactites duriaei            GGAAGCGCACTCCAAATACCTCAATG-TGTTGTCTCTTGATGACGTTTCG
Cynara humilis                GGAAGCGCTCTCCATAGACCCCAACG-TGTCGTCCTGTGACGACACTTCG
Lamyropsis cynaroides         GGAAGCGCTCTCCAAAGACCCCAACG-TGTCGTCTTGTGACGACGCTTCG
 
                                       510       520       530       540        
character                      .........|.........|.........|.........|.........
gap character                     0000000001111111111222222222233333333334444444
                                  1234567890123456789012345678901234567890123456
Ancathia igniaria              ACC0000010001100001?00?????0001000100000100010010
Olgaea pectinata               ACC0000010010100000000000000001001000000100010000
Synurus deltoides              ACC0000010000100001000000000001000010000000010000
Alfredia cernua                ???0000010010000001??????????????????????????????
Onopordum boissieri            ACC0000010100100001000000000001000000100000010000
Carlina vulgaris               ACC0100?10010100111000000001000010100000000011000
Xeranthemum cylindraceum       ACC0000010101100101010101100000010100000000010000
Echinops exaltatus             ACC0100011001100001010101100001100100000100010000
Berardia subacaulis            ACC0001010001110001000000100000010100000000010100
Stemmacantha rhapontica        ACC0010110010100001001010100000010010000010110000
Serratula coronata             ACC0010110001100001010000000000010010000010110000
Klasea radiata                 ACC0010110001100001000000100000010010000010110100
Stephanochilus omphalodes      ACC0010110001000001011001100001000010000010110000
Jurinea macrocephala           ACC0000010001000001000000000011100100000010100000
Jurinea mollis                 ACC0000010001000000000000000011100110000111100000
Saussurea riederi              ACC0000010001000001000000000000010010000010110000
Cousinia hystrix               ACC000001000100000110000000000001001000001011000?
Arctium lappa                  ACC0000010001111000110010001000010011000011110000
Carduus crispus                ACC0000000001100001000100010001010010000000000010
Carduus nutans                 ACC0000000001100001000100010001011010000000011010
Carduus defloratus             ACC0000000001100001000100010001010110010000010000
Cirsium tuberosum              ACC0000000001100001000000100001010010000000010001
Cirsium texanum                ACC0001001001100011000010000001010010000000010000
Tyrimnus leucographus          ACC0000000001100001000000100100010010000000010000
Cirsium ciliatum               ACC0000000001100001000000100101010110000000010000
Cirsium eriophorum             ACC0000000001100001000000100000010110000000010111
Notobasis syriaca              ACC0000000001100001000000000001010010110000010000
Carduus leptacanthus           ACC0000000001100001000000000001010010000000010000
Ptilostemon afer               ACC1000000001100001000000000000010010000000010000
Galactites duriaei             ACC1000000001101000000000000000010011000000010000
Cynara humilis                 ACC0000000001111011000000001000010011001000010000
Lamyropsis cynaroides          ACC0000000001100001000000100001000010001000010000