Prof. Dr. Thomas Borsch

Directorate

Director

phone

+49 30 838 50133

fax

+49 30 838-4 50133

direktor(at)bgbm.org

Botanischer Garten und Botanisches Museum Berlin-Dahlem

Freie Universität Berlin

Königin-Luise-Straße 6-8

14195

Berlin

Deutschland

Tasks

Director of the Botanic Garden and the Botanical Museum Berlin-Dahlem, Freie Universität Berlin

Professor (W3) at the Institute of Biology, Systematic Botany and Plants Geography at the Freie Universität Berlin

Curriculum vitae

married, 2 daughters
born 18 February 1969 in Königstein im Taunus
Studies in Biology 1992-1996 in Frankfurt and Bonn
Dr. rer. nat. in Biology in 2000, Universität Bonn

Scientific career / Employments

1992-1994	Research Assistant (part-time), Senckenberg Research Institute, Frankfurt
1999-2000	Research Assistant, Institute of Botany, Universität Bonn
2000-2005	Scientific Assistant (C1), Nees Institute, Universität Bonn
2005-2007	Privatdozent, Nees Institute, Head of the laboratories of molecular systematics
2007-2008	Professor (W2) for Biodiversity and Evolution of plants, Carl von Ossietzky Universität Oldenburg, Director of the Botanic Garden of the Universität Oldenburg
2008-	Professor (W3) at the Institute of Biology, Systematic Botany and Plants Geography at the Freie Universität Berlin and Director of the Botanic Garden and the Botanical Museum Berlin-Dahlem, Freie Universität Berlin

Stays abroad

Visiting scientist at the Virginia Polytechnic Institute and State University, USA (1997-1999), Visiting scientist at the Royal Botanic Gardens Kew (1990, 6 months), at the Missouri Botanical Garden (1992, 3 months), and research stays (1-3 months) in Ethiopia, Bolivia, Mexico and Venezuela.

Research interests

Evolution of flowering plants; molecular phylogenetics and especially mutational dynamics on non-coding genomic regions and their application of molecular markers; pollen morphology and ultrastructure research; genesis of biological diversity (based on the study of model groups such as the plant families Amaranthaceae and Nymphaeaceae); Assessment of plant diversity and taxonomy; endemism and biogeography of plants in biodiversity hotspots (especially Cuba and the Caucasus); conservation and sustainable use of biodiversity.

Honours and awards

2007- Referee, Commission for Botany, Akademie der Literatur und der Wissenschaften, Mainz

2006-2007 Scholarship Heisenberg, Deutsche Forschungsgemeinschaft (DFG)

1997-1999 Scholarship Studienstiftung des deutschen Volkes

1996 Heinrich-Hörlein Gedächtnispreis, Universität Bonn

1987 “Jugend forscht” 2. Bundespreis in Biologie

Editorships

Journal of Systematics and Evolution (Associate Editor): 2009 - 2014

Plant Systematics and Evolution (Special Volume Editor, with D. Quandt and M. Koch): 2008

Revista Mexicana de Biodiversidad (Editoral Advisory Board): 2007-

TAXON (Special Volume Editor, with P.S. Soltis): 2008

Willdenowia (Editoral Board): 2008 -

Mandates in scientific advisory boards and committees

Netherlands Center for Biodiversity Naturalis, Scientific Advisory Board

Consortium of European Taxonomic Facilities (CETAF), BoD

European Distributed Institute of Taxonomy (EDIT), BoD

Dahlem Center of Plant Sciences (DCPS), Deputy Speaker

Memberships in scientific societies

American Society of Plant Taxonomists

Botanical Society of America

Deutsche Botanische Gesellschaft

Gesellschaft für Biologische Systematik

International Association for Plant Taxonomy

International Organisation of Systematics and Evolutionary Biology (President 2009-2011)

Sociedad Mexicana de Botánica

Sociedad Cubana de Botánica (corresponding member)

weitere Aktivitäten

Organizing activities

2006	17th Symposium of Systematics and Biodiversity of the German Botanical Society; Meeting organiser
2011	Biosystematics 2011, International Congress of Systematics and Evolutionary Biology; Congress President

Publications

Papers in peer reviewed journals (published and in press)

91.	Canal D., Köster N., Korotkova N., Jones K.E., Croat T.B. & Borsch T. in press: Phylogeny and diversification history of the large neotropical genus Philodendron (Araceae): Accelerated speciation in a lineage dominated by epiphytes. – Amer. J. Bot.
90.	Di Vincenzo V., Gruenstaeudl M., Nauheimer L., Wondafrash M., Kamau P., Demissew S. & Borsch T. 2018: Evolutionary diversification of the African achyranthoid clade (Amaranthaceae) in the context of sterile flower evolution and epizoochory. – Ann. Bot. dor.org/10.1093/aob/mcy055
89.	Duwe V.K., Muller L.A.H., Reichel K., Zippel E., Borsch T. & Ismail S.A. 2018: Genetic structure and genetic diversity of the endangered grassland plant Crepis mollis (Jacq.) Asch. as a basis for conservation management in Germany. - Conserv. Genet., doi 10.1007/s10592-017-1025-8
88.	Torres-Montúfar A., Borsch T. & Ochoterena H. 2018: When homoplasy is not homoplasy: Dissecting trait evolution by contrasting composite and reductive coding. - Syst. Biol. 67: 543-551, doi: 10.1093/sysbio/syx053
87.	Ismail S.A., Duwe V.K., Zippel E. & Borsch T. 2018: Assessment of current genetic structure from local to geographic scales indicates brake down of historically extensive gene flow in the dry grassland species Scabiosa canescens Waldst. & Kit. (Dipsacaceae). - Divers. Distr. 24: 233-243, doi: 10.1111/ddi/12667
86.	Korotkova N., Parolly G., Khachatryan A., Ghulikyan L., Akopian J. A., Borsch T. & Gruenstaeudl M. 2018: Towards resolving the evolutionary history of Caucasian pears (Pyrus, Rosaceae) - Phylogenetic relationships, divergence times and leaf trait evolution. – J. Syst. Evol. 56: 35-47, doi: 10.111/jse.12276
85.	Maharramova E., Gruenstaeudl M., Huseynova Irade M., Kolbaia S., Borsch T. & Muller L. A. H. 2018: Phylogeography and population genetics of the riparian relict tree Pterocarya fraxinifolia (Juglandaceae) in the South Caucasus – Syst. Biodiv. 16: 14-27, doi: 10.1080/147720000.2017.1333540
84.	Torres-Montúfar A., Borsch T., Fuentes S., Clase T., Peguero B. & Ochoterena H. 2017: The new Hispaniolan genus Tainus (Rubiaceae) constitutes an isolated lineage in the Caribbean biodiversity hotspot. Willdenowia 47: 259-270, doi 10.3327/wi.47.47309
83.	Korotkova N., Borsch T. & Arias S. 2017: A phylogenetic framework for the Hylocereeae (Cactaceae) and implications for the circumscription of the genera. Phytotaxa 327: 1-46, doi: 10.11646/phytotaxa.327.1.1
82.	Gruenstaeudl M., Nauheimer L. & Borsch T. 2017: Plastid genome structure and phylogenomics of Nymphaeales: Conserved gene order and new insights into relationships. - Plant Syst. Evol. 303: 1251-1270, doi: 10.1007/s00606-017-1436-5
81.	Malekmohammadi M., Akhani H. & Borsch T. 2017: Phylogenetic relationships of Limonium (Plumbaginaceae) inferred from multiple chloroplast and nuclear loci. - Taxon 66: 1128-1146, doi: 10.12705/665.8
80.	Jones K., Korotkova N., Petersen J., Henning T., Borsch T. & Kilian N. 2017: Dynamic diversification history with rate upshifts in Holarctic bell-flowers (Campanula and allies). – Cladistics 33: 637-666, doi:10.1111/cla.12187
79.	Duwe V.K., Muller L.A.H., Borsch T. & Ismail S.A. 2017: Pervasive genetic differentiation among Central European populations of the threatened Arnica montana L. and genetic erosion at lower elevations. - Perspectives Pl. Ecol. Evol. Syst. 27: 45-56, dx.doi.org/10.1016/j.ppees.2017.02.003
78.	Lücking R., Dal Forno M., Moncada B., Coca L.F., Vargas-Mendoza L.Y., Aptroot A., Arias J.L., Besal B., Bungartz F., Cabrera-Amaya D.M. & al. 2016: Turbo-taxonomy to assemble a megadiverse lichen genus: seventy new species of Cora (Basidiomycota: Agaricales: Hygrophoraceae), honouring David Leslie Hawksworth’s seventieth birthday. Fungal Div. doi: 10.1007/s13225-016-0374-9
77.	Cervantes A., Fuentes S., Gutierrez J., Magallón S. & Borsch T. 2016: Successive arrivals since the Miocene shaped the diversity of the Caribbean Acalyphoideae (Euphorbiaceae) – J. Biogeogr. 43: 1773-1785, doi:10.1111/jbi.12790
76.	Duwe V. K., Muller L. A. H., Borsch T. & Ismail S. A. 2016: Development of microsatellite for Crepis mollis (Astaraceae) – Applic. Pl. Sci. 4 (7), doi:10.3732/apps.1600022
75.	Geiger M. F., Astrin J.J., Borsch T., Burkhardt U., Grobe P., Hand R., Hausmann A., Hohberg K., Krogmann L., Lutz M., Monje C., Misof B., Morinière J., Müller K., Pietsch S., Quandt D., Rulik B., Scholler M., Traunspurger W., Haszprunar G. & Wägele W. 2016: How to tackle the molecular species inventory for an industrialized nation – lessons from the first phase of the German Barcode of Life initiative GBOL – Genome 59: 1-10, http://dx.doi.org/10.1139/gen-2015-0185
74.	Robson D. B., Wiersema J. H., Hellquist C. B. & Borsch T. 2016: Distribution and Ecology of a New Species of Water-lily, Nymphaea loriana (Nymphaeaceae), in Western Canada. – Canad. Field-Naturalist 130: 25–31.
73.	Flores-Olvera H., Zumaya S., Borsch T. 2016: Two new species of Iresine (Amaranthaceae: Gomphrenoideae) from Mexico supported by morphological and molecular characters. – Willdenowia 46: 165-174. doi: http://dxdoi.org/10.3372/wi.46.46113
72.	Kilian N., Henning T., Plitzner P., Müller A., Güntsch A., Stöver B. C., Müller K. F., Berendsohn W. G., Borsch T. 2015: Sample data processing in an additive and reproducible taxonomic workflow by using character data persistently linked to preserved individual specimens. – Database 2015: bav094 http:/dx.doi.org/10.1093/database/bav094
71.	Hernández-Ledesma P., Berendsohn W. G., Borsch T., Mering S. von, Akhani H., Arias S., Castañeda-Noa I., Eggli U., Eriksson R., Flores-Olvera H., Fuentes-Bazán S., Kadereit G., Klak C., Korotkova N., Nyffeler R., Ocampo G., Ochoterena H., Oxelman B., Rabeler R. K., Sanchez A., Schlumpberger B. O. & Uotila P. 2015: A taxonomic backbone for the global synthesis of species diversity in the angiosperm order Caryophyllales - Willdenowia 45: 281 – 384. http://dx.doi.org/10.3372/wi.45.45(link is external) 301
70.	Borsch T., Hernández-Ledesma P., Berendsohn W. G., Flores-Olvera H., Ochoterena H., Zuloaga F. O., von Mering S. & Kilian N. 2015: An integrative and dynamic approach for monographing species-rich plant groups - building the global synthesis of the angiosperm order Caryophyllales. – Perspectives Pl. Ecol. Evol. Syst. 17(4), 284–300. doi:10.1016/j.ppees.2015.05.003
69.	Rillig M.C., Kiessling W., Borsch T., Gessler A., Greenwood A.D., Hofer H., Joshi J., Schröder B., Thonicke K., Tockner K., Weisshuhn K. & Jeltsch F. 2015: Biodiversity research: data without theory – theory without data. Front. Ecol. Evol. 3:20. Doi: 10.3389/fevo.2015.00020

68.	Ismail S.A., Duwe V.K., Fleischer-Notter H., Borsch T. 2015: Development of ten polymorphic microsatellite markers for Scabiosa canescens Waldst. & Kit. (Dipsacaceae). – Conservation Genet Resour 7:725-728. doi:10.1007/s12686-015-0446-2
67.	Duwe V. K., Ismail S. A., Buser A., Sossai E., Borsch T., & Muller L. A. H. 2015: Fourteen Polymorphic Microsatellite Markers for the Threatened Arnica montana (Asteraceae). – Appl. Plant Sci. 3(1). http://dx.doi.org/10.3732/apps.1400091
66.	Maharramova E., Safarov H., Kozlowski G., Borsch T. & Muller L. A. 2015: Analysis of nuclear microsattelites reveals limited differentiation between Colchic and Hyrcanian populations of the wind-pollinated relict tree Zelkova carpinifolia. - American Journal of Botany 102 (1): 119 – 128. doi: 10.3732/ajb.1400370
65.	Borsch T. & Löhne C. 2014: Botanic Gardens for the future: Integrating research conservation, environmental education and public recreation – Ethiop. J. Biol. Sci. 13 (Suppl.): 115–133.
64.	Korotkova N., Nauheimer L., Ter-Voskayan H., Allgaier M. & Borsch T. 2014: Variability among the most rapidly evolving plastid genomic regions is lineage-specific: implications of pairwise genome comparisons in Pyrus (Rosaceae) and other angiosperms for marker choice. - PLoS One 9 (11): e112998. doi: http://dx.doi.org/10.1371/journal.pone.0112998
63.	Borsch T., Wiersema J. H., Hellquist C. B., Löhne C. & Govers K. 2014: Speciation in North American water-lilies: Evidence for the hybrid origin of the newly discovered Canadian endemic Nymphaea loriana (Nymphaeaceae) in a past contact zone. - Canadian Journal of Botany 92 (12): 867–882. doi.org/10.1139/cjb-2014-0060
62.	Crowl A. A., Mavrodiev E., Mansion G., Haberle R., Pistarino A., Kamari G., Phitos D., Borsch T. & Cellinese N. 2014: Phylogeny of Campanuloideae (Campanulaceae) with emphasis on the utility of nuclear pentatricopeptide repeat (PPR) genes. - PLoS ONE 9 (4): e94199. doi:10.1371/journal.pone.0094199
61.	Maharramova E. H., Muller L. H., Korotkova N. & Borsch T. 2014: Development of nuclear microsatellites for the Arcto-Tertiary tree Zelkova carpinifolia (Ulmaceae) using 454 pyrosequencing. – Appl. Plant Sci. 2 (3). doi:103732/apps.1300072
60.	Tesfaye K., Govers K., Bekele E. & Borsch, T. 2014: ISSR fingerprinting of Coffea arabica throughout Ethiopia reveals high variability in wild populations and distinguishes them from landraces. – Plant Syst. Evol. 300 (5): 881-897. doi: http://dx.doi.org/10.1007/s00606-013-0927-2
59.	Bécquer E., Michelangeli F.A. & Borsch T. 2014: Comparative seed morphology of the Antillean genus Calycogonium (Melastomataceae: Miconieae) as a source of characters to untangle its complex taxonomy. - Phytotaxa 166 (4): 241–258. http://dx.doi.org/10.11646/phytotaxa.166.4.1
58.	Meneses R. I., Borsch T., Ortuño T. & Fuentes A. 2013: Hacia una flora de Bolivia. – Rev. Soc. Boliv. Bot. 7: 97-105.
57.	González Gutiérrez P. A., Köhler E. & Borsch T. 2013: New species of Buxus (Buxaceae) from northeastern Cuba based on morphological and molecular characters, including some comments on molecular diagnosis – Willdenowia 43: 125–137. http://dx.doi.org/10.3372/wi.43.43115
56.	Zumaya S., Flores-Olvera, H. & Borsch T. 2013: Two new Mexican endemic species of Iresine (Amaranthaceae) – Syst. Bot. 38(2):434-443. doi: http://dx.doi.org/10.1600/036364413X666633
55.	Mansion G., Parolly G., Crowl A.A., Mavrodiev E., Cellinese N., Oganesian M., Fraunhofer K., Kamari G., Phitos D., Haberle R., Akaydin G., Ikinci N., Raus T. & Borsch T. 2012: How to handle speciose clades? Massive taxon sampling as a strategy towards illuminating the natural history of Campanula (Campanuloideae). – PLoS ONE 7(11): e50076. doi:10.1371/journal.pone.0050076
54.	Fuentes Bazan S., Uotila P. & Borsch T. 2012: A novel phylogeny-based generic classification for Chenopodium sensu lato, and a tribal rearrangement of Chenopodioideae (Chenopodiaceae). – Willdenowia 42: 5-24. doi: 10.3372/wi42.42101
53.	Sánchez-del Pino Y., Motley T.J. & Borsch T. 2012: Molecular phylogenetics of Alternanthera (Gomphrenoideae, Amaranthaceae): resolving a complex taxonomic history caused by different interpretations of morphological charactersin a lineage with C4 and C3-C4 intermediate species. – Bot. J. Linn. Soc. 169: 493-517.
52.	Barniske A., Borsch T., Müller K., Krug M., Neinhuis C. & Quandt D. 2012: Phylogenetics of early branching eudicots: comparing phylogenetic signal across chloroplast introns, spacers and genes. – J. Syst. Evol. 50: 85-108. doi: 10.1111/j.1759-6831. 2012.00181.x
51.	Fuentes S., Mansion G. & Borsch T. 2012: Towards a species level tree of the globally diverse genus Chenopodium (Chenopodiaceae). – Molec. Phylogenet. Evol. 62: 359-374. doi: 10.1016/j.ympev.2011.10.006.
50.	Korotkova N., Borsch T., Quandt D., Taylor N.P., Müller K.F. & Barthlott W. 2011: What does it take to resolve relationships and to identify species with molecular markers? An example from the epiphytic Rhipsalideae (Cactaceae). – Amer. J. Bot. 98: 1549-1572. doi:10.3732/ajb.1000502.
49.	Borsch T., Ortuño Limarino T. & Nee M. H. 2011: Phylogenetics of the neotropical liana genus Pedersenia (Amaranthaceae: Gomphrenoideae) and discovery of a new species from Bolivia based on molecules and morphology. – Willdenowia 41: 5 – 14. doi:10.3372/wi.41.41101.
48.	Košuth J., Smelcerovic A., Borsch T., Zuehlke S., Karppinen K., Spiteller M., Hohtola A. & Čellárová E. 2011: The hyp-1 gene is not a limiting factor for hypericin biosynthesis in the genus Hypericum – Funct. Pl. Biol. 38: 35-43. doi: 10.1071/FP10144.
47.	Borsch T., Löhne C., Mbaye M.S. & Wiersema J. 2010: Towards a complete species tree of Nymphaea: shedding further light on subg. Brachyceras and its relationships to the Australian waterlilies – Telopea 13: 193-217.
46.	Schäferhoff B., Fleischmann A., Fischer E., Albach D. C., Borsch T., Heubl G., & Müller K. F.: Towards resolving Lamiales relationships: insights from rapidly evolving chloroplast sequences. – BMC Evol. Biol. 10: 352. doi: 10. 1186/1471-2148-10-352.
45.	Löhne C., Wiersema J. H. & Borsch T. 2009: The unusual Ondinea, actually just another Australian water-lily of Nymphaea subg. Anecphya (Nymphaeaceae). – Willdenowia 39: 55-58.
44.	Schäferhoff B., Müller K. F. & Borsch T. 2009: Caryophyllales phylogenetics: disentangling Phytolaccaceae and Molluginaceae and description of Microteaceae as a new isolated family. – Willdenowia 39: 209-228.
43.	Korotkova N., Schneider J. V., Quandt D., Worberg A., Zizka G. & Borsch T. 2009: Phylogeny of the eudicot order Malpighiales. Analysis of a recalcitrant clade with sequences of the petD group II intron. – Pl. Syst. Evol. 282 : 201-228.
42.	Borsch T. & Quandt D. 2009: Mutational dynamics and phylogenetic utility of non-coding chloroplast DNA. – Pl. Syst. Evol. 282 : 169-199.
41.	Borsch T., Korotkova N., Raus T., Lobin W. & Löhne C. 2009: The petD group II intron as a species level marker: utility for tree inference and species identification in the diverse genus Campanula (Campanulaceae). – Willdenowia 39: 7-33.
40.	Kusari S., Zühlke S., Borsch T. & Spiteller M. 2009: Positive correlations between hypericin and putative precursors detected in the quantitative secondary metabolite spectrum of Hypericum. – Phytochemistry 70: 1222-1232.
39.	Worberg A., Alford M. H., Quandt D. & Borsch T. 2009: Huerteales sister to Brassicales plus Malvales, and newly circumscribed to include Dipentodon, Gerrardina, Huertea, Perrotetia and Tapiscia. – Taxon 58: 468-478.
38.	Sánchez del-Pino I., Borsch T. & Motley T. 2009: trnL-F and rpl16 sequence data and dense taxon sampling reveal monophyly of unilocular anthered Gomphrenoideae (Amaranthaceae) and an improved picture of their internal relationships. – Syst. Bot. 34: 57-67.
37.	Gole T.W., Borsch T., Denich M. & Teketay D. 2008: Floristic composition and environmental factors characterizing coffee forests in southwest Ethiopia. – Forest Ecol. Managem. 255: 2138-2150.
36.	Borsch T. & Soltis P. S. 2008: Nymphaeales – the first globally diverse clade? – Taxon 57: 1051.
35.	Borsch T., Wiersema J. H. & Löhne C. 2008: Phylogeny and evolutionary patterns in Nymphaeales: integrating genes, genomes and morphology. – Taxon 57: 1052-1081.
34.	Löhne C., Yoo M.-Y., Borsch T., Bell C.D., Wiersema J. H., Wilde V., Soltis P. S., Soltis D. E. & Barthlott W. 2008: Biogeography of Nymphaeales – extant patterns and historical events. – Taxon 57: 1123-1146.
33.	Löhne C., Borsch T., Jacobs S. W. L., Hellquist C. B., Wiersema J. H. 2008: Nuclear and plastid DNA sequences reveal complex reticulate patterns in Australian water lilies (Nymphaea subgenus Anecphya, Nymphaeaceae). – Austral. Syst. Bot. 21: 229-250.
32.	Beck S. G., Fleischmann A., Huaylla H., Müller K. F. & Borsch T. 2008: Pinguicula chuquisacensis (Lentibulariaceae), a new species from the Bolivian Andes, and first insights on phylogenetic relationships among South American Pinguicula. – Willdenowia 38: 201-212.
31.	Sage R., Sage T.L., Pearcy, R.W., Borsch T. 2007: The taxonomic distribution of C4 photosynthesis in Amaranthaceae sensu stricto. – Amer. J. Bot. 94: 1992-2003.
30.	Tesfaye G. K., Borsch T., Govers K., Bekele E. 2007: Characterisation of Coffea chloroplast microsatellites and evidence for the recent divergence of C. arabica and C. eugenioides cp genomes. – Genome 50: 1112-1129.
29.	Löhne C., Borsch T. & Wiersema JH. 2007: Phylogenetic analysis of Nymphaeales using fast-evolving and non-coding chloroplast markers. – Bot. J. Linn. Soc. 154: 141-163.
28.	Borsch T., Hilu KW., Wiersema JH., Löhne C., Barthlott W. & Wilde V. 2007: Phylogeny of Nymphaea (Nymphaeaceae): evidence from substitutions and microstructural changes in the chloroplast trnT-trnF region. – Int. J. Pl. Sci. 168: 639-671.
27.	Worberg A, Quandt D., Barniske A-M., Löhne C., Hilu KW. & Borsch T. 2007: Phylogeny of basal eudicots: insights from non-coding and rapidly evolving DNA. – Organisms Diversity Evol. 7: 55-77.
26.	Wanke S, Jaramillo MA., Borsch T., Samain SM., Quandt D., Neinhuis C. 2007: Evolution of Piperales – matK gene and trnK intron sequence data reveal lineage specific resolution contrast. – Molec. Phylogenet. Evol. 42: 477-497.
25.	Ortuño T. & Borsch T. 2006: A further new species of Gomphrena (Amaranthaceae, Gomphrenoideae) from the dry valleys of Bolivia. – Kew Bull. 61: 565-568.
24.	Müller K., Borsch T., Legendre L., Porembski S. & Barthlott W. 2006: Recent progress in understanding the evolution of Lentibulariaceae. – Pl. Biol. 8: 748-757.
23.	Greilhuber J., Borsch T., Müller M., Worberg A., Porembski S. & Barthlott W. 2006: Smallest angiosperm genomes found in Lentibulariaceae, with chromosomes of bacterial size. – Pl. Biol. 8: 770-777.
22.	Müller KF., Borsch T. & Hilu KW. 2006: Phylogenetic utility of rapidly evolving DNA at high taxonomical levels: contrasting matK, trnT-F and rbcL in basal angiosperms. – Molec. Phylogenet. Evol. 41: 99-117.
21.	Moore S E M., Hemsley A R. & Borsch T. 2006: Micromorphology of outer exospore coatings in Selaginella megaspores. – Grana 45: 9-21.
20.	Müller K. & Borsch T. 2005: Stellate pore ornamentation in Amaranthaceae pollen: Multiple origins of a unique feature. – Grana 44: 266-281.
19.	Cieslack T., Polepalli J.S., White A., Müller K., Borsch T., Barthlott W., Steiger J., Marchant A. & Legendre L. 2005: Phylogenetic analysis of Pinguicula (Lentibulariaceae): chloroplast DNA sequences and morphology reveal several geographically distinct radiations. – Amer. J. Bot. 92: 1723-1736.
18.	Qiu Y-L, Dombrovska O., Lee J., Li L., Whitlock B.A., Bernasconi-Quadroni F., Rest J.S., Borsch T., Hilu K.W., Renner S., Soltis D.E., Soltis P.S., Zanis M.J., Gutell J. C., R., Powell M., Savolainen V., Chatrou L.W. & Chase M.W. 2005: Phylogenetic analyses of basal angiosperms based on nine plastid, mitochondrial, and nuclear genes. – Int. J. Pl. Sci. 166: 815-842.
17.	Woods K., Hilu K.W., Borsch T. & Wiersema J.H., 2005: Pattern of variation and systematics of Nymphaea odorata: II. Sequence information from ITS and trnL-trnF. – Syst. Bot. 30: 481-493.
16.	Woods K., Hilu K.W., Wiersema J.H. & Borsch T. 2005: Pattern of variation and systematics of Nymphaea odorata: I. Evidence from morphology and Inter-Simple Sequence Repeats (ISSRs). – Syst. Bot. 30: 471-480.
15.	Müller K. & Borsch T. 2005: Phylogenetics of Amaranthaceae using matK/trnK sequence data – evidence from parsimony, likelihood and Bayesian approaches. – Ann. Missouri Bot. Gard. 92: 66-102.
14.	Rahmanzadeh R., Müller K., Fischer E., Bartels E. & Borsch T. 2005: Linderniaceae and Gratiolaceae are further lineages distinct from Scrophulariaceae (Lamiales). – Pl. Biol. 7: 67-78.
13.	Neinhuis C., Wanke S., Hilu K.W., Müller K. & Borsch T. 2005: Phylogeny of Aristolochiaceae based on parsimony, likelihood, and Bayesian analyses of trnL-trnF sequences. – Pl. Syst. Evol. 250: 7-26.
12.	Löhne C. & Borsch T. 2005: Phylogenetic utility and molecular evolution of the petD group II intron in basal angiosperms. – Molec. Biol. Evol. 22: 317-332.
11.	Müller K. & Borsch T. 2005: Phylogenetics of Utricularia (Lentibulariaceae) and molecular evolution of the trnK intron in a lineage with high mutational rates. – Pl. Syst. Evol. 250: 39-67.
10.	Ortuño T. & Borsch T. 2005: Dos nuevas especies de Gomphrena (Amaranthaceae, Gomphrenoideae) de los valles secos de Bolivia. – Novon 15: 180-189.
9.	Müller K., Borsch T., Legendre L., Porembski S., Theisen I. & Barthlott W. 2004: Evolution of carnivory in Lamiales and Lentibulariaceae. – Pl. Biol. 6: 477- 490.
8.	Kadereit G, Borsch T., Weising K. & Freitag H. 2003: Phylogeny of Amaranthaceae and Chenopodiaceae and the evolution of C4 photosynthesis. – Int. J. Pl. Sci. 164: 959-986.
7.	Hilu KW., Borsch T., Müller K., Soltis D.S., Soltis P.S., Savolainen V., Chase M.W., Powell M., Alice L.A., Evans R., Campbell C., Sauquet H., Neinhuis C., Slotta T., Rohwer J. & Chatrou L. 2003: Angiosperm phylogeny based on matK sequence information. – Amer. J. Bot. 90: 1758-1776.
6.	Sauquet H., Doyle J.A., Scharaschkin T., Borsch T., Hilu K.W., Chatrou L.W., Thomas A. Le. 2003: Phylogenetic analysis of Magnoliales and Myristicaceae based on multiple data sets: implications for character evolution. – Bot. J. Linn. Soc. 142: 125-186.
5.	Borsch T., Hilu K.W., Quandt D., Wilde V., Neinhuis C. & Barthlott W. 2003: Non-coding plastid trnT-trnF sequences reveal a well resolved phylogeny of basal angiosperms. – J. Evol. Biol. 16: 558-576.
4.	Borsch T., Clemants S. & Mosyakin S. 2001: Symposium: Biology of the Amaranthaceae-Chenopodiaceae-alliance. – J. Torrey Bot. Soc. 128: 234- 235.
3.	Borsch T. 1998: Pollen types in the Amaranthaceae. Morphology and evolutionary significance. – Grana 37: 129-142.
2.	Borsch T. & Barthlott W. 1998: Structure and evolution of metareticulate pollen. – Grana 37: 68-78.
1.	Borsch T. 1995: Three new combinations in Pfaffia (Amaranthaceae) from the New World tropics. – Novon 5: 230-233.

Book chapters and monographs (published and in press)

10.	Borsch T., Ortuño T. & Nee M. (in press): Amaranthaceae. . – Monogr. Syst. Bot. Missouri Bot. Gard.
9.	Chase M.W., Fay M.F., Devey D.S., RØnsted N., Davies J., Pillon Y., Petersen G., Seberg O., Tamura M.N., Asmussen C.B., Hilu K.W., Borsch T., Davis J.L., Stevenson D.W., Pires J.C., Givnish T.J., Sytsma K.J. & Graham S.W. 2006: Multi-gene analyses of monocot relationships: a summary. – Pp. 63-75 in: Columbus J.T. (ed.), Proceedings on the third international conference on the monocots. – Aliso 22
8.	Quandt D., Müller K., Stech M., Hilu K.W., Frey W., Frahm J.P. & Borsch T. 2004: Molecular evolution of the chloroplast trnL-F region in land plants. – Pp. 13-37 in: Goffinet B., Hollowell V., and Magill R. (ed.), Molecular systematics of bryophytes. – Monogr. Syst. Bot. Missouri Bot. Gard. 98.
7.	Bojan B., Clemants S. & Borsch T. 2003: Amaranthaceae. – Pp. 415-429 in: ZY Wu & PR Raven (ed.), Flora of China 5. – Beijing: Science Press & St Louis: Missouri Botanical Garden.
6.	Barthlott W., Theisen I., Borsch T. & Neinhuis C. 2003: Epicuticular waxes and vascular plant systematics: integrating micromorphological and chemical data. – Pp. 189-206 in: Stuessy T.F., Mayer V., and Hörandl E. (ed.), Deep morphology: toward a renaissance of morphology in plant systematics. – Regnum Veg. 141.
5.	Borsch T. 2001: Amaranthaceae. – Pp. 56-83 in: Stevens W.D., Ulloa Ulloa C., Pool A. & Montiel O. (ed.), Flora de Nicaragua 1. – Monogr. Syst. Bot. Missouri Bot. Gard. 80.
4.	Borsch T. & Wilde V. 2000: Pollen variability within species, populations, and individuals, with particular reference to Nelumbo nucifera. – Pp. 285-299 in: Harley M., Blackmore S. & Morton C. (ed.), Pollen and spores: morphology and biology. Kew: Royal Botanic Gardens.
3.	Borsch T. & Wolter M. 1998: Pollenmorphologie und -ultrastruktur. – Pp. 135-145 in: Barthlott W. & Lobin W. (ed.), Amorphophallus titanum. – Trop. Subtrop. Pflanzenwelt 99.
2.	Borsch T., Neinhuis C. & Barthlott W. 1998: Nelumbo: new data on an exceptional plant. – Pp. 408-416 in: Auluo Z. & Sugong W. (ed.), Floristic characteristics and diversity of East Asian plants. – Kunming: China Higher Education Press & Hamburg: Springer.
1.	Borsch T. 1993: Amaranthaceae. – Pp. 18-26 in: Brako L. & Zarucchi J.L. (ed.), Catalogue of the flowering plants and gymnosperms of Peru / Catálogo de las angiospermas y gymnospermas del Peru. – Monogr. Syst. Bot. Missouri Bot. Gard. 45.

Journals, not peer reviewed

6.	Borsch T., Löhne C., Müller M., Hilu K.W., Wanke S., Worberg A., Barthlott W., Neinhuis C. & Quandt D. 2005: Towards understanding basal angiosperm diversification: recent insights using rapidly evolving genomic regions. – Nova Acta Leopoldina, ser. 2, 92: 85-110.
5.	Borsch T. & Pedersen T.M. 1997: Restoring the generic rank of Hebanthe Martius (Amaranthaceae). – Sendtnera 4: 13-31.
4.	Borsch T. & Barthlott W. 1995: Classification and distribution of the genus Nelumbo (Nelumbonaceae). – Beitr. Biol. Pfl. 68: 421-450.
3.	Borsch T. & Kühn K. 1995: Bodenparameter und die standörtliche Differenzierung bachbegleitender Vegetation. Eine Studie im Vortaunus. – Courier Forschungsinst. Senckenberg 186 (Hans Joachim Conert-Festschrift): 169-183.
2.	Borsch T. 1993: Das Gartenportrait: Der Missouri Botanical Garden. – Palmengarten 57: 26-31.
1.	Borsch T. 1990: Die Vegetation extensiv genutzten und brachliegenden Grünlands im Naturschutzgebiet „Hinterste Neuwiese“ (Vortaunus). Untersuchungen zur Sukzession auf Feuchtwiesen. – Bot. Naturschutz Hessen 4: 14-45.

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Prof. Dr. Thomas Borsch

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Botanischer Garten und Botanisches Museum Berlin